22 
SPERG U LARI A 
In addition to such hereditary variations, the plant, of course, varies a great deal in response to the situation in which 
it grows : in the shelter of larger plants and in wet mud, large individuals occur with comparatively long internodes, whilst 
in drier and exposed situations the individuals are prostrate, small, and the internodes very short. There is no reliable 
evidence that variations of this kind, at least among the higher plants, are inherited. 
The inland stations of this species are of considerable ecological interest. In Berkshire, the plant occurs in a flat 
marshy meadow along with the following species : — Tolypella glomerata, Sagina nodosa , Ranunculus sceleratus, Apiurn graveolens, 
Scirpus compressus, S. maritimus, and Car ex distans. Mr G. C. Druce (FI. Berksh. 103) thinks that the plant was conveyed 
to this place by birds, and that its continued existence there is due to the saline nature of the habitat. 
In Worcestershire, the plant occurs by the side of the Droitwich Canal. “ How it got there is a matter of speculation : 
probably the seeds were brought there through the agency of natural dispersion, and finding a congenial locality, owing to 
the saline conditions of the place, have reproduced their kind.” The plant occurs also on Defford Common “where a 
salt spring once existed.” Other halophytes or hemihalophytes occurring in saline habitats of Worcestershire are : — Atriplex 
glabriuscula var. babingtoni, Glaux 7 naritima, Glyceria maritima , and Juncus compressus var. gerardi. See Amphlett and Rea 
The Botany of Worcestershire (1909), p. 59 et passim. 
In Cheshire, the species also occurs in inland localities, as “ by canals and roads, etc., particularly in the salt district ” 
of Northwich and Winsford (de Tabley in FI. Cheshire 54 (1899)). 
Not uncommon on the drier edges of salt-marshes in nearly all the maritime counties of 
the British Islands ; rare on the wetter parts of salt-marshes ; northwards to Orkney ; very local 
in brackish inland localities, as in Berkshire, Worcestershire, and Cheshire. 
Scandinavia, Denmark, Germany, Holland, Belgium, France, central Europe, Russia, southern 
Europe ; northern Africa ; Asia ; America. 
S', marginata x salina (see below); S. rupicola x salina (p. 19). 
Series ii. Marinae 
Marinae nobis. For characters, see page 18. 
British species and hybrid of Marinae 
S. marginata x salina (see below). Capsules (when fertile) intermediate in size between those of the 
putative parents. Seeds usually dimorphic, some wingless and others winged, rarely all with very narrow wings 
or none at all. 
5. S. marginata (p. 23). Capsules markedly longer than the calyx. Seeds suborbicular, 
winged. 
S. marginata x salina hybr. nov. ; Arenaria glandulosa Jacquin Hort. Schoenbr. iii, 56 (1798); Lepi- 
gonum glandulosum Kindberg Monogr. Lepig. 14 et 16, fig. 2 (1863); S. dillenii Rouy et Foucaud FI. France iii, 
303 (1896) excl. race salina , et excl. syn. L. medium Fries et L. leiospermum Kindberg. 
leones : — More in Eng. Bot. Suppl. t. 2978, as Lepigonum salinum (repeated in Syme Eng. Bot. t. 255, as 
S. neglecta)\ Jacquin Hort. Schondr. t. 355, as A renaria glandulosa. 
Camb. Brit. FI. iii. Plate 20. (a) Portion of flowering shoot, (b) Portion of shoot with stipules and two 
laminae (enlarged). ( c ) Calyx with barren capsule (enlarged). ( d ) Calyx with fertile capsule (enlarged). 
(e) Seeds (enlarged). Isle of Wight (E. W. H.). 
Exsiccata : — Lojacono {PI. Sic. Par.), as Lepigonum heterospermum. 
Perennial, often a very much larger plant than either S', salina or S', marginata. Shoot 
often strongly glandular, especially on the younger portions. Branches usually prostrate, some- 
what compressed, slightly 2-ribbed. Stipules comparatively smaller, narrower, and more acute 
than in S. salina. Laminae nearly terete, thicker and longer than in S. salina. Inflorescence 
more elongate than in S. salina and with more flowers. Pedicel of the lower flowers a little 
longer than the persistent calyx. Flowers about 1 cm. in diameter ; May to August. Sepals 
about as broad and as long as the petals. Petals rose-purple, with a paler almost white base, 
a little larger than in S. salina. Stamens usually 10, obdiplostemonous. Capsules often of two 
kinds ; some about as long as the persistent calyx and containing many abortive seeds, others 
a little longer than the calyx and containing fertile seeds. Seeds dimorphic, mostly wingless and 
pyriform, a few suborbicular and surrounded by a membranous and radially marked wing, all 
with a more or less thickened rim, smooth or nearly so. 
Merely glandular forms of A. marginata should not be referred to S. marginata x salina. 
Possibly the two following plants should also be referred to this putative hybrid : — S. marginata var. angustata Clavaud 
FI. Gironde in Act. Soc. Linn. Bordeaux xxxv, 403 (1881) and Lepigonum marinum var. apterum Marshall in Journ. Bot. xxxix, 
268 (1901) which resemble S. marginata in habit, but have shorter capsules and the wing of the seed rudimentary or absent. 
Rare, on salt-marshes and spray-washed rocks; Jersey, Isle of Wight, Somerset, and possibly elsewhere. 
Scandinavia, Denmark, France ; Africa ; and probably elsewhere. 
