SPERGULARIA 
23 
5. SPERGULARIA MARGINATA. Large-flowered Sea Spurrey. Plates 21 ; 20 
Spergula marina Dalechamp Hist. Plant. 1385 (1586); cf. Johnson in Gerard Herball ed. 2, 1125 (1633); 
Alsine spergula major maritima jlore violacea Morison Hist, ii, 551 (1680); Spergula marina nostras Ray Hist. 
1034 (1688); Alsine spergula facie media C. Bauhin Pinax 251 (1671); Ray Syn. ed. 3, 351 (1724); Arenaria 
foliis linearibus longitudine internodiorum L. Hort. Cliff. 173 (1737). 
Spergularia marginata Kittel Taschenb. ed. 2, 1003 (1844); Syme Eng. Bot. ii, 13 1 (1864) excl. t. 
25 7; Rouy et Foucaud FI. France iii, 302 (1896); Arenaria rubra var. marina L. loc. cit ., parti m ; Ar. media 
Linn, herb., non Sp. PI. ed. 2, 606 (1762); DC. Prodr. iii, 422 (1822); Wallroth Sched. Crit. 202 (1822); 
nomen confusum ; Ar. marina Allioni FI. Pedem. ii, 114 (1785) partim ; Roth loc. cit., partim ; Withering Arr. iii, 
422 (1796) partim; Persoon Syn. 504 (1805); non Wallroth; nomen confusum; Ar. marginata DC. FI. France 
iv, 793 (1805); Lepigo7ium marinum Wahlberg FI. Gothob. 47 (1820) descr. gen. nulla; Fries FI. Suec. Mant. iii, 
35 (1842); Kindberg Syn. Lepig. 12 (1856); Monogr. Lepig. 14 et 18, fig. 6 (1863) excl. syn. Cambessides ; Ar. 
marina var. succosior Mertens und Koch Deutschl. FI. iii, 294 (1821); Alsine marina Wahlenberg FI. Suec. i, 281 
(1824); Hiern in fourn. Bot. xxxvii, 318(1899); excl. syn. Ar. media L. ; non Reichenbach ; S. media C. B. Presl 
FI. Sic. i, 161 (1826) excl. syn. L. ; t Buda marina Dumortier FI. Belg. no (1827) nomen; Alsine 77iarginata 
Reichenbach FI. Germ. Excurs. 566 (1830); Al. marma var. obesior Koch Syn. iii (1835); Lepigonum mar- 
ginatum Koch in Flora xxi, 505 (1841). 
leones: — Smith Eng. Bot. t. 958, as Are7iaria marina ; Svensk Bot. t. 743, as Alsine 77iarina ; DC. Icon. 
t. 48, as Arenaria marginata. 
Ca7nb. Brit. FI. iii. Plate 21. (a) Flowering shoot. ( b ) Seeds (enlarged). Norfolk (C. E. M.). ( c ) Flowering 
shoot. ( d ) Seed (enlarged). Somerset (E. W. H.). ( e ) Flowering shoot. (/) Seed (enlarged). Isle of 
Wight (E. W. H.). 
Exsiccata : — Bourgeau {PI. d'Esp.), 975 ; Fries, viii, 38, as Lepig 07 iu 7 n marinum ; Wirtgen, viii, 330, as 
Lepig 07 ium marginatum. 
Perennial, a larger plant than S. salina. Shoot glandular or eglandular. Branches prostrate 
or decumbent, more or less compressed, terete. Stipules smaller than in S. salina and S. mar- 
ginata x salina , entire except sometimes in age. Laminae larger than in S. salina. fnflorescence 
rarely dichasial. Pedicel of the lower flowers markedly longer than the persistent calyx, some- 
times about twice as long. Flowers about i'2 — 1 - 5 cm. in diameter, appearing earlier than those 
of S. salina ; mid-May to September. Sepals narrower than and a little shorter than the petals. 
Petals pale lilac with a white base, much paler in colour and markedly larger than those of 
■S', salina. Stamens 10, obdiplostemonous. Capsules larger than in S. salina and 5 . marginata 
x salina, projecting considerably from the calyx, about 10—12 mm. long and broad, acute. Seeds 
broadly pyriform to suborbicular, compressed, usually rimmed, all surrounded by a membranous 
radially marked wing. 
The Arenaria rubra var. marina L. includes both Spergularia salina and S. marginata, as seen particularly in Linnaeus’s 
FI. Atiglica (1754). The Arenaria media L. Sp. PL (1762) does not refer to either of those species, but to Spergula 
pentandra, though the specimen of the Linnaean herbarium is Spergularia marginata and a specimen by Linnaeus at Stock- 
holm is {fide Kindberg) 5. rupicola. 
We definitely reject the trivial names media and marina for this species and the last as nomitia confusa, and take up 
the unequivocal names salina and marginata. 
(J 3 ) subvar. glandulosa comb. nov. ; Buda media var. glandulosa Druce in Bot. Exch. Club Brit. Rep. for 
1899, i, p. 599 (1901)! 
Shoot more or less strongly glandular. 
Kent (Druce loc. cit.) and doubtless elsewhere. 
Abundant on salt-marshes (especially in the general or mixed salt-marsh association), occasional 
on spray-washed rocks, rather rare on the drier edges of salt-marshes (where it sometimes grows 
side by side with S. salina) ; in nearly all the maritime counties of the British Islands, north- 
wards to Orkney ; unknown in inland localities. 
Scandinavia, Denmark, Germany, Holland, Belgium, France, central Europe, Russia, southern 
Europe ; Africa ; Asia ; America. 
S', marginata x salina (page 22). 
Tribe III. SAGINEAE 
Sagineae Caruel in Ledebour’s 
PI. 963 (1836 — 1840). 
For characters, see page 14. 
FI. Ital. ix, 564 (1892) non Fenzl ; Sabulmeae Fenzl in Endlicher Ge?i. 
