VACCARIA 
85 
leones: — Smith Eng. Bot. t. 1577; Reichenbach Icon, vi, t. 302. 
Camb. Brit. FI. iii. Plate 81. ( a ) Flowering shoot. (b) Flowers. (c) Ovary. Hort. (E. M. H.). 
Exsiccata : — Billot, 1432 ; Bourgeau (PI. Lyc.), 38 ; Dickson, iv, 9 (hort.) ; Gandoger (FI. Gall. Exsicc.), 
11 6, as C. sphaericus', 999, as C. lugdunensis ; Lange, 365; Wirtgen, xvi, 886. 
Shoot pubescent. Branches 4 — 6 dm., diffuse. Petioles short or almost absent. Laminae oval, 
acute, up to about 4 cm. long and 2 broad. Inflorescence 3 — 4-flowered, dichasial above. Flowers 
July and August. Calyx green or purplish brown, with 5 sepals. Petals tinged with green or yellow, 
deeply bifid. Styles 3. Fruit globular, about 8 — 10 mm. in diameter, succulent, ultimately black 
and dry. Seeds black, shining, punctate. 
Very rare and not indigenous; on the banks of a ditch in the Isle of Dogs, Middlesex, “but almost 
certainly introduced ” (Syme op. cit. 54), and now extinct there ; adventitious elsewhere, as in Norfolk. 
Germany, Holland, France, central Europe, Russia, southern Europe; Asia. 
Tribe VII. DIANTHEAE 
Diantheae Reichenbach Handb. Natiirl. Pflanzen. 298 (1837); A. Braun in Flora xxvi, 377 (1843); Pax 
op. cit. 69 et 74 (1889); Rouy et Foucaud FI. France iii, 150 (1896). 
For characters, see page 14. 
British genera of Diantheae 
Genus 17. # Vaccaria (see below). Epicalyx absent. Calyx 5-angular, not umbilicate. Petals 
with claw and limb gradually passing into each other ; coronal scales absent. 
Genus 18. fSaponaria (p. 86). Epicalyx absent. Calyx terete, umbilicate. Petals with limb 
and claw well defined ; coronal scales absent. 
Genus 19. Tunica (p. 86). Epicalyx present, loose-fitting, soon becoming scarious, covering 
all or most of the calyx. Petals with limb and claw gradually passing into each other ; coronal 
scales absent. 
Genus 20. Dianthus (p. 88). Epicalyx present, tight-fitting, herbaceous during the flowering 
period, covering the lower half or less than half of the calyx. Petals with limb and claw well-defined ; 
coronal scales present. 
Genus 17. * Vaccaria 
Vaccaria Moench Meth. PI. 63 (1794); Al. Braun in Flora xxvi, 381 (1843); Willkomm et Lange 
Prodr. FI. Hisp. iii, 672 (1878); Pax op. cit. 75 et 76 (1889); Rouy et Foucaud FI. France iii, 155 (1896). 
Epicalyx absent. Calyx oval-pyramidal, 5-angled, many-veined, not umbilicate. Gynophore very 
short. Petals with claw and limb passing gradually into each other, claw winged, coronal scales absent. 
Stamens 5 + 5. Capsule opening by 4 short teeth, persistently biseptate towards the base. Embryo 
horse-shoe-shaped. Seeds reniform to subglobose. 
The genus Vaccaria is often cited as of Medicus Phil. Bot. i, 96 (1789); but no such genus is there established. 
3 species ; Europe ; western Asia. 
[1. # VACCARIA PARVIFLORA. Plate 82] 
Vaccaria Gerard Herball 395 (1597). 
Vaccaria parviflora Moench Meth. PI. 63 (1794); Rouy et Foucaud FI. France iii, 155 (1896); Saponaria 
vaccaria L. Sp. PI. 409 (1753) ; Miller Gard. Diet. ed. 8, no. 3 (1768); V. pyramidata Medicus Phil. Bot. i, 96 (1789) 
descr. gen. nulla; 5. segetalis Necker Delic. Gallo-Belg. 194 (1768) nomen abortivum ; Lychnis vaccaria Scopoli FI. 
Cam. ed. 2, ii, 303 (1772); S'. rubra Lamarck FI. Fr. ii, 541 (1778) nomen abortivum; Gypsoplnla vaccaria 
Sibthorp et Smith FI. Graec. Prodr. i, 279 (1806). 
leones : — Reichenbach Icon, vi, t. 245. 
Camb. Brit. FI. iii. Plate 82. (a) Flowering branch, (b) Petal, (c) Ovary. Hort., origin Nottinghamshire. 
Exsiccata : — Billot, 728, as Gypsophila vaccaria ; Reichenbach, 2499, as V. pyramidata ; Thielens et Devos, 
i, 39, as S', vaccaria. 
Annual. Shoot glabrous, somewhat glaucous. Stem erect, 3 — 6 dm., leafy. Leaves — lower ones 
oblong, attenuate at the base ; upper ones sessile, lanceolate, subconnate, cordate, acuminate. Pedicels 
long. Flowers about 15 cm, in diameter; June and July. Calyx not umbilicate, pyramidal. Petals 
rose-pink; limb short, Capsule ovoid, shorter than the calyx. Seeds black, large. 
P 
