I IO 
AQUILEGIA 
Subtribe III. A Q UILEGIINA E 
Aquilegiinae nobis; Isopyreae Syme Eng. Bot. i, 59 (1863). 
For characters, see page 104. Only British genus: — Aquilegia. 
Genus 5. Aquilegia 
Aquilegia [Tournefort Inst. 428, t. 242 (1700);] L. Sp. PI. 533 (1753) et Gen. PI. ed. 5, 237 (1754); 
Prantl op. cit. 56 et 59 (1891). 
Perennial. Laminae 2-ternately or 3-ternately compound. Flowers actinomorphic. Sepals 5, 
petaloid, deciduous. Petals or nectar-leaves 5, large, coloured, each with a spreading tip and a long 
basal curved hollow spur which passes backwards between the sepals and then curves round towards 
the axis. Stamens in about 10 whorls of 5 each; inner ones sterile, with dilated filaments appressed 
to the ovary. Styles slender. Follicles 5, slightly united at the base, erect, sessile. Seeds biseriate. 
About 50 species ; northern hemisphere. 
I. AQUILEGIA VULGARIS. Columbine. Plate 108 
A. coendea Gerard Herball 935 (1597), inch A. rubra et A. multiplex', A. flore simplici Ray Syn. ed. 3, 273 
(1724). 
Aquilegia vulgaris L. Sp. PI. 533(1753); Hudson FI. Angl. 207 (1762), inch A. alpina p.208; Smith 
Eng. Bot. no. 297 (1796); FI. Brit. 578 (1800); Syme Eng. Bot. i, 60 (1863); Rouy et Foucaud FI. France i, 
123 (1893). 
Map 44. Distribution of Aquilegia vulgaris in the British Islands 
leones: — Smith Eng. Bot. t. 297; FI. Dan. t. 695; Svensk Bot. t. 118; Reichenbach Icon, iv, t. 114, fig. 
4729 ; Syme Eng. Bot. i, t. 46. 
Camb. Brit. FI. iii. Plate 108. (a) Lower Leaf. ( b ) Flowering branch. ( c ) Fruiting branch. ( d ) Follicle. 
(e) Follicle in transverse section. (/) Seeds. Cumberland (L. B.). 
Exsiccata : — Billot, 1407; Reichenbach, 1092, as A . nigricans ; 1289; 1590, as A. vulgaris var. platysepala ; 
Todaro, 1207; Herb. FI. Ingric. ix, 25. 
