CORY DA LIS 
1 68 
Family 2. FUMARIACEAE 
Fumariaceae DC. Syst. Nat. ii, 105 (1821); Battling Ordines Nat. PI. 259 (1830); Bernhardi in Linnaea 
viii, 465 (1833); Rouy et Foucaud FI. France i, 169 (1893); Fwnario'ideae Al. Braun in Ascherson FI. Brandenb. 
i, 48 (1864); Prantl und Kiindig in Pfianzenfam. iii, pt. ii, 137 et 142 (1889). 
Perennial or annual herbs, usually glabrous, destitute of latex. Leaves spirally arranged (i.e., 
“alternate”), much divided; leaflets stalked. Inflorescence racemose, terminal or leaf-opposed. 
Bracteoles 2, median. Flowers transversely zygomorphic, pedicelled. Sepals 2, generally caducous, 
originally median. Petals 4 ; 2 of them outer, larger, and originally lateral ; the other 2 inner, smaller, 
and originally median : the orientation becomes apparently reversed owing to a twist of the pedicel 
through an angle of 90° ; the apparently upper petal prolonged backwards into a swollen nectar- 
containing spur ; outer petals winged, more or less coherent before insect-visitation. Stamens 2, 
originally lateral, each more or less separated at maturity into 3 branches, the middle staminal branch 
bearing a bilocular anther, the lateral branches with a unilocular half-anther, usually emerging (some- 
times, as in C. capno'ides , explosively) on insect-visitation, the lateral stamens (as well as the lateral 
sepals) suppressed, a staminal outgrowth protecting the nectar. Ovary superior, of 2 carpels, unilocular. 
Ovules 2, anatropous. Fruit with few or 2 or 1 seeds. Placentation parietal. Embryo minute. 
Cotyledons linear. Endospei r m oily. 
7 genera, and about 150 species; extra-tropical northern hemisphere, South Africa. 
British genera of FUMARIACEAE 
Genus 1. Corydalis (see below). Ovules 00. Fruit a capsule, dehiscing by the two carpels, 
elongate, strongly compressed laterally. Seeds with an aril. 
Genus 2. Fumaria (p. 1 7 1 ). Ovules 2. Fruit indehiscent, 1 -seeded, subglobose. Seeds 
without an aril. 
Genus 1. Corydalis 
Corydalis [Dillenius Cat. PI. Giss. app. Tab. 7 (1718) ;] Ventenat Choix 19(1803); DC. FI. France iv, 636 
(1805); Prantl und Kiindig in Pfianzenfam. iii, pt. 2, 143 et 144 (1891); nomen conservatum ; non Medicus 
Phil. Bot. i, 96 (1789); Fumaria L. loc. cit., pro max. parte; Capno'ides [Tournefort Inst. 423, t. 237 (1700); 
Adanson Fain. PI. ii, 431 (1763) incl. Cisticapnos ;] Neckera Scopoli Introd. Hist. Nat. 313 (1 777); Pseudo- 
Fumaria [Rivinus] Medicus Phil. Bot. i, 110(1789). 
Perennial or annual herbs. Allied to Fumaria, but differing in the following characters. 
Flowers usually larger and more variedly coloured. Ovules 00 . Style more or less persistent. 
Fruit a capsule, dehiscing by the two carpels, elongate, bilaterally compressed. Seeds arillate. 
About 90 species ; Europe, Asia, Africa. 
Sections of Corydalis 
Section I. # Capnites (see below). Subterranean tuber present. Inflorescence terminal. 
Section II. Capno’ides (p. 169). Rhizome slender, tuber not present. Inflorescence lateral. 
Section I. *CA PNITES 
Capnites DC. Syst. Nat. ii, 115 (1821); Bulbocapnos Bernhardi in Linnaea vii, 604 (1832) as a genus; 
Rouy et Foucaud FI France i, 184 (1893). 
For characters, see above. Only British species: — *C. bulbosa. 
i. ^CORYDALIS BULBOSA. Plate 177 
Corydalis bulbosa DC. FI France iv, 637(1805); N. E. Brown in Eng. Bot. ed. 3, suppl. 21 (1891); 
Fumaria bulbosa var. solida L. Sp. PI. 699 (1753); F. bulbosa Miller Gat'd. Diet. ed. 8, no. 8 (1768); F. solida 
Ehrhart Bcitr. vi, 146(1791); Smith FI. Brit. 748(1800); F. intermedia Withering Bot. Arr. ed. 3, iii, 620, t. 29 
