Danaea simp licif alia, Rudge. 
1 1 5 
Cotyledon. 
The adult cotyledon is a very simple spathulate leaf 
without stipules 1 (cf. Figs. 3 and 4), and the venation is 
monopodial in its branching, thus agreeing with Angiopteris 2 
and differing from Marctttia Douglasii 3 . 
The vascular bundle is very simple, consisting sometimes 
of only two or three tracheids and a small amount of phloem. 
It may be considered collateral, but this depends on whether 
the layer of parenchymatous cells towards the upper surface 
is to be looked upon as phloem or otherwise. Certain it is 
that phloem, with recognizable sieve-tubes, occurs only on 
the underside and flanks of the bundle. 
The Stem. 
The primary stem, which is comparatively short, has a small 
concentric bundle, or stele, of the ordinary Marattiaceous type, 
and is somewhat elongated in the same plane as the diarch 
xylem-plate of the root, the stele of which merges so gradually 
into that of the stem that it is impossible to determine where 
the one ends and the other begins. With regard to the 
phloem, the transition first shows itself by a gradual increase 
in the number of the thin-walled elements at the poles of the 
xylem-plate of the root, so that the protoxylem no longer 
abuts directly on the endodermis, or is separated from it by 
of embryos have been examined under conditions more favourable for determining 
their orientation. 
1 Danaea agrees with the other genera of the Marattiaceae in having well- 
developed stipules to the older fronds, and, moreover, the sclerenchyma of the 
petiole, &c. is of the usual Marattiaceous type. Holle in his paper ‘ Ueber die 
Vegetationsorgane der Marattiaceen (Sitzung der Koniglichen Gesellschaft der 
Wissenschaften zu Gottingen, Jan. 8, 1876), made a mistake in these two respects, 
which was subsequently corrected by Kuhn ( Ueber den anatomischen Bau von 
Danaea , Flora, 1890), who pointed out that Danaea differs anatomically in no 
essential respect from the rest of the Marattiaceae. There is little doubt, as 
Kuhn suggests, that Holle had incorrectly-named specimens to deal with. I myself 
have had the opportunity of verifying Kuhn’s statements in several species of 
Danaea. 
2 Farmer, loc. cit., Fig. 19. 
3 Campbell, loc. cit. (a), p. 11 and Fig. 27. 
