196 Phillips. — On the Development of the 
As I have shown, this is a misunderstanding of the nature of 
these filaments ; the term ‘ carpogenous system ’ is a misnomer 
as applied to them, for they are sterile, and the real sporogenous 
filaments arise later, not from their terminal cells, but below 
from the auxiliary cell. The period of rest which Janczewski 
found to exist may, however, admit of explanation. I have 
frequently observed cystocarps of the outward shape of 
mature cystocarps, but with only the tufts of sterile filaments 
occupying a small space at the base of the cavity. It has 
occurred to me that these are cystocarps in which the con- 
jugation of the carpogonium with the auxiliary cell has either 
failed to take place or been greatly delayed. Otherwise the 
repose is nothing more than the cessation in the growth of 
the sterile filaments. The function of these sterile branches 
is partly nutritive perhaps, but they probably also serve the 
purpose of opening out a cavity for the cystocarp in the dense 
tissue, by means of their mucilaginous cell-walls. They have, # 
however, with a varying degree of development, a remarkable 
morphological constancy in all the Rhodomelaceae which 
I have examined. 
Askenasy (12) seems more recently to have described the 
development of the cystocarp in Dasya Berkeleyi (Mont.), J. Ag., 
and found that the phenomena were similar to what occurred 
in Polysiphonia . Dasya Berkeleyi has, however, latterly been 
removed by Agardh (10 b) from Dasya to the genus Hetero- 
siphonia , Mont. This course is also followed by Schmitz (8 a). 
But while the plant might therefore be expected to depart 
somewhat from the normal type of Dasya, still, inasmuch as 
it presents the character of a sympodial growth like Dasya , it 
might be expected in the structure of its cystocarp to show 
affinities to Dasya rather than to Polysiphonia. Apparently 
this is not the case. I regret I have not been able to consult 
Askenasy’s paper directly. 
In the same work (10) by Agardh as that in the course of 
which he has re-established the genus Heterosiphonia , Mont., he 
has generally revised the genus Dasya , and discussed the struc- 
ture of the ‘ nucleus,’ in the genus as reconstituted. He refers to 
