Cystocarp in Rhodomelaceae (//). 197 
the difference in the appearance of the 4 nucleus ’ when it is 
less evolved, and consists of shorter, slightly-branched fila- 
ments, from that of the more fully-developed f nucleus ’ with 
greatly-prolonged and much-branched filaments. This differ- 
ence, I do not doubt, is the difference between the cystocarp 
when it contains only tufts of sterile filaments, and when 
it contains the larger system of sporiferous filaments. Writing 
of Dasya coccinea itself (10 a), he says : c In Dasya coccinea 
cuius nucleum fasciculo basali simpliciore constitutum obser- 
vavi, iam dignoscere licet plures ramos inferne firmiores, quos 
sensim magis evolutos fieri facilius assumeres.’ These £ rami 
firmiores ’ are almost certainly the sterile earlier contents of 
the cystocarp, and, as my figures show, they do not gradually 
develop into the sporiferous threads. 
Chondria tenuissima, C. A g. 
Here, as in Rhodomela and Polysiphonia , I find the procarp 
is formed upon one of the so-called leaves, and upon the second 
joint from the axis. Considering that the axis is relatively 
stout and strongly corticated, it might seem unlikely that only 
one stalk-cell intervened between the procarp and the axis. 
This is, however, the case (Fig. 8), as the cell becomes greatly 
elongated and attenuated. Janczewski’s figures (6 a) of this 
plant err in showing more than one axial cell in the pedicel. 
Bornet and Thuret’s figure (13), however, makes the point 
clear. 
At the time of the fertilization of the trichogyne, the two 
sterile branches have already attained much the same degree 
of development as that which they reach long after this stage 
in Dasya. The paranemata have not yet arisen from the 
central cell, and the four-celled carpogonial branch lies in 
a lateral position, pushed on one side by the luxuriant growth 
of the lateral sterile branch. After fertilization the sterile 
filaments cease to grow almost entirely. The paranemata, 
however, grow apace, and the upper portion of the cystocarp is 
formed almost wholly by means of these rows of cells. The 
