348 
Burt • — The Development of 
system of the mycelial strand ; M, the continuation of the 
medullary system of that strand ; and N, the upper end of M — 
come to play very unequal parts in later development. Of 
these parts C becomes the outermost layer of the volva ; M , 
the bundle of medullary tissue connecting the mycelial strand 
with the base of the stipe ; while the sheaf-like head N gives 
rise to all the remaining parts, including stipe, gleba, and 
gelatinous layer of the volva. 
In eggs of this stage stained in toto in Delafield’s haema- 
toxylin or in P. Mayer’s haemalum, the hyphae of the 
medullary bundle were about 4 \x in diameter. They had 
very thin walls and were divided by extremely thin cross- 
walls into cells about 20 /x long. In most of these cells 
there could be seen a deeply-stained, opaque, spherical mass 
about 2 ju. in diameter — presumably the nucleus. In the sheaf- 
like head N, the tissue was so intricate from the crossing of 
hyphae in different planes that the cross-septa could not be 
made out with certainty. The nuclei were, however, very 
numerous in this portion and not more than 3 or 4 /x apart. 
The hyphae of the cortical portion are from 4 to 5 in 
diameter and from 20 to 25 /x long. 
Ed. Fischer 1 described as young a stage of Ithyphallus im- 
pudicus , shown in his Fig. 18, which differs from my Fig. 1 chiefly 
in the greater number of hyphal branches from the medullary 
bundle (M of my Fig. 1) passing out nearly at right angles 
into the cortical layer. In having more such branches in the 
cortical layer of its egg than exist in M. caninus , the cortical 
tissue of the egg /. impudicus merely shares a character of the 
cortical system of its mycelial strand, as the latter has been 
made known by De Bary 2 . On account of the presence of 
such branches in the egg of I. impudicus , Fischer separated 
the tissue which I have designated by C into an outer peri- 
pheral zone and an inner mass bordering on the medullary 
bundle. The inner mass he supposed to be identical with the 
1 L. c., 1890, p. 22. 
2 Zur Morphologic der Phalloideen. In Beitrage z. Morph, u. Physiol, der Pilze, 
Abhandl. d. Senkenb. Naturf.-Ges., Bd. V, 1864, p. 204. 
