354 Burt . — The Development of 
In his recent paper on M. caninus , Ed. Fischer has pointed 
out the connexion between the masses of tramal and inter- 
mediate tissue, and has concluded that the gleba must originate 
from the differentiation of portions of the intermediate tissue 1 . 
The course of development does not favour this conclusion 
(see Fig. 4), but shows rather that the intermediate tissue 
A and that of the gleba, although both arising from the sheaf- 
like head N, by the differentiation of originally intimately 
connected portions, tend to almost wholly lose such con- 
nexions through developmental changes, and have nothing 
further in common in their development. In such further 
development the glebal tissue, making its tramal connexion 
with the upper end of the central column R of apparently 
increasing functional importance, takes on the formation of 
spores; while the intermediate tissue A, losing also its con- 
nexions with the sides of the column R, but retaining those 
with the basal portions of the egg, goes on to form a recepta- 
culum of varied form for the different genera — a supporting 
and elevating structure for the spore-mass. 
The rudiment of the wall of the stipe is in the stage of 
Fig. 5, a narrow layer, S, of swollen portions of hyphae very 
irregular in form and very intimately connected with the loose 
intermediate tissue A, of which they form a part. The hyphae 
of this rudiment lie close against, and often interlock with, 
the longitudinally running hyphae of the column R. The 
details of this structure are shown in Fig. 7, which represents 
under higher magnification a portion of the rudiment of the 
stipe with adjacent tissues at the lower end of the stipe. The 
exact position of this area in Fig. 5 is indicated by 5 . The 
hyphae on the left in Fig. 7 belong to the tissue of the central 
column R ; those on the right are in the intermediate tissue 
A ; the more deeply shaded hyphae between constitute the 
rudiment of the stipe 5 . The septa of the hyphae did not 
show distinctly in the preparation, and no attempt at their 
representation has been made in the figure. The hyphae of 
1 L. c., 1895, p. 136. 
