362 Burt. — The Development of 
receptaculum, and for the dense shell of pseudoparenchyma 
about each chamber, and next to the central column R , and 
next to the intermediate tissue, by stating — 
1. That in early development the tissue of the rudiment 
of the stipe differentiates into a series of isolated dense hyphal 
knots, which are separated from the central column R } from 
each other, and from the tissue A, by narrow open spaces ; 
2. That an outgrowth of swollen hyphal ends — pseudoparen- 
chyma — then takes place into these spaces from the isolated 
knots of hyphae, from the central column R, and from the 
tissue A ; 
3. That these approaching plates of pseudoparenchyma 
become loosely grown together later ; 
4. That the central portions of the hyphal knots become 
constantly looser and more gelatinous, finally forming the 
chamber-cavities of the wall. 
There are important features in the above that need modi- 
fication. It seems probable that Fischer’s hyphal knots are 
the portions c, c, Fig. 1 1, which stain lighter in my prepara- 
tions ; but these portions are certainly not isolated from each 
other by open spaces as he states and figures 1 . On the 
contrary, the light-staining portions are closely connected 
together from their first appearance by a densely interwoven 
tissue of which they are only portions. These connexions are 
retained in the various preparations which I have seen. It is 
quite true that branches from the tissue of the light portions — 
the chambers later — crowd their way into the plates and 
become pseudoparenchyma, thus increasing the area of these 
plates : pseudoparenchymatous hyphae, favourably situated in 
the partition-wall, also branch and help to augment the same 
surfaces. 
Nature of the pits' in the ‘spore-bearing' part of the 
wall. — In the mature plant pits are found opening from the 
wall of the stipe into its main central cavity. These pits are 
in the upper portion of the stipe — in its spore-bearing portion. 
1 L, c., 1887, figs. 13, 17, 18. 
