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PACIFIC SCIENCE, Vol. XI, April, 1957 
for a person who is familiar with the species 
to determine, in the majority of cases, from 
which one of the islands a specimen was 
collected. However, this broad generalization 
of geographical distribution is not constant, 
and almost all forms are present on each of the 
islands where the species occurs. 
Of the naturalized species Eugenia Cumini 
is by far the most widely distributed. E. 
malaccensis is confined mostly to shaded, 
moist valleys where it is likely they were 
planted originally by the Hawaiians. Both 
Eugenia Jambos and E. uniflora are only sparsely 
distributed in the native forests. 
These introduced species are remarkably 
constant in their characters and show no sig- 
nificant degree of variation. 
EXPLANATIONS 
Unless otherwise indicated, the specimens 
studied are in the Bernice P. Bishop Museum. 
The standard abbreviations of Lanjouw and 
Stafleu (1954) are used to indicate the loca- 
tions of the other specimens: 
BISH — Bernice P. Bishop Museum, 
Honolulu 
GH— Gray Herbarium, Cambridge 
K — Royal Botanical Gardens, Kew 
M — Botanische Staatssammlung, 
Miinchen 
MICH— University of Michigan, Ann Arbor 
NY — New York Botanical Gardens, 
New York 
US — United States National Herbarium, 
Washington 
I have examined the following numbers of 
fruits and seeds of the species included in this 
problem. When possible both fresh and dried 
material was studied. 
Fresh Dried 
Eugenia malaccensis 15 3 
E. Jambos — 5 
E. sandwicensis 100 60 
E. Cumini 50 20 
E. uniflora 10 5 
E. rariflora — 15 
E. koolauensis 20 10 
TAXONOMY 
Eugenia L., Sp. PL 470, 1753. 
Syzygium Gaertn., Fruct. 1: 166, t. 33, 1788. 
Jambosa DC., Prodr. 3: 286, 1828. 
Trees or shrubs. Shoots glabrous or pube- 
scent. Leaves simple, opposite, glandular- 
punctate, pinnately veined with a continuous 
intramarginal vein. Leaf scar with a single 
vascular bundle. Flowers single or in pairs, 
axillary, or in terminal, or axillary cymes or 
racemes, or inflorescence on leafless branches. 
Calyx tube 5 globose to elongate-turbinate, 
extending beyond the ovary or not so, usually 
minutely glandular-punctate; calyx lobes 4, 
large persistent and spreading, or small and 
early deciduous; petals 4, free and spreading 
persistent or caducous, or cohering and falling 
off as a calyptra; stamens numerous, free on a 
staminal disk lining the calyx tube or inserted 
on the margin of the calyx tube; anthers versa- 
tile, cells splitting longitudinally, connective 
gland present; style filiform, stigma small; 
ovary inferior, 2-celled. Fruit a berry with 
only 1 seed (rarely 2) developing from the 
many ovules, crowned by the persistent calyx 
lobes or by the truncate scars of the calyx 
lobes; umbilicus sometimes present; seeds 
large with a thin membranaceous or a thick 
cartilaginous or fibrous seed coat; cotyledons 
thick, fleshy, completely free or partly or 
entirely fused. 
Type species: Eugenia uniflora L. 
KEY TO THE SPECIES IN HAWAII 
A. Flowers in cymes or racemes; calyx tube 
extending beyond the ovary. 
B. Calyx tube 10-30 mm. long. 
C. Inflorescence axillary from older 
leafless nodes or rarely from leafy 
5 "Calyx tube” is used here, as is customary in 
treatments of the genus, to refer to the inferior ovary 
and especially to the tissues adhering to it. There is no 
intention to imply the exact morphological nature of 
the inferior ovary of Eugenia by the use of this term. 
The origin of the inferior ovary in this genus has not 
as yet been satisfactorily determined. 
