174 
PACIFIC SCIENCE, Vol. XI, April, 1957 
(M); U. S. Explor., Exped., highland near 
Waianae; large branch of Lualualei Valley, 
southwest of Pohakea Pass, Aug. 4, 1932, 
Degener and Bush 7,299 (NY) . 
No locality: Collect. Dr. Hillebrand, Oahu, 
Maui (GH). 
Molokai: Central Molokai, wet forest, 
Oct. 13, 1916, Hitchcock 13,193 (US). 
Maui: Olowalu Valley, May 19, 1920, 
Forbes 2417. M (US); Wailuku, W. Maui, Hil- 
lebrand and Lydgate. 
Eugenia rariflora was first recorded for the 
Hawaiian Islands by Hillebrand in 1888. It 
occurs occasionally in the drier regions of the 
larger islands except Hawaii. This species 
shows considerable variation in its leaves al- 
though not to as great a degree as E . sand- 
wicensis. 
Both E. waianensis and E. koolauensis var. 
glabra are here reduced to Eugenia rariflora 
because of intermediate forms which make it 
impossible to set definite limits upon the two. 
The concave leaves would be the only char- 
acter which would separate E. koolauensis var. 
glabra from E. rariflora. But a collection from 
Palikea, Oahu ( Degener 7,303) has leaves 
which are only slightly concave; another spec- 
imen ( Degener 20,381) shows leaves that are 
more strongly concave. This character is cer- 
tainly not a reliable one, and is of no taxo- 
nomic value in this case. This conclusion is 
supported by the fact that the characters of 
E. koolauensis var. glabra are identical in all 
other respects with those of E. rariflora. 
The size of the leaves of E. rariflora might 
seem to offer a remarkably valuable taxonomic 
character, particularly when the extremes in 
form are considered. E. waianensis is described 
as differing from E. rariflora in having smaller 
leaves. This character seems to be of value 
when specimens with extremely small leaves 
are studied ( Degener 7,296). However, some 
specimens, such as those collected by Forbes 
(2417. M) and Rock (17,003), have leaves 
which show a series of intermediate sizes be- 
tween E. waianensis and E. rariflora. In such 
cases this character proves to be of little value. 
7. Eugenia koolauensis Deg., FI. Hawaii. 
Fam. 273, 8/10/32. 
Tree 4-7 m. tall; branches grey, glabrate, 
in age the bark reddish-brown, black spotted; 
leaf scars 1-2 mm. wide, rounded shield- 
shaped, reddish-brown; leafy branchlets 1-3 
mm. in diameter, 4-angled to terete, densely 
brown subappressed-pilosulous; internodes 
9-19 mm. long; leaves 2.5-5 cm. long, 1-3.3 
cm. wide, obovate to elliptic, apex obtuse or 
apiculate, base subcuneate, blade coriaceous, 
concave, margin entire, more or less strongly 
revolute, above olive-green, minutely glandu- 
lar-punctate, glabrous and shiny or subap- 
pressed-pilosulous near veins with inter- 
vals glabrous, below pale yellowish-green, 
minutely pitted or pustulate, subappressed- 
puberulent; midrib shallowly impressed above, 
elevated below; primary lateral veins, alter- 
nate or opposite, 5-7 on a side, 5-10 mm. 
apart, irregularly ascending at 140-150°, 
meeting in an irregularly lobed intramarginal 
vein 1-2 mm. from leaf margin, raised on 
both surfaces but more distinct below, rarely 
slightly shallowly impressed above; the vein- 
lets raised-reticulate; petioles 2-4 mm. long, 
1 mm. wide, brown subappressed-pilosulous; 
flowers single or in pairs, axillary; peduncles 
1-8 mm. long, subappressed-pilosulous, brac- 
teate; bract subulate, yellow-brown subap- 
pressed-pilosulous, 1 mm. long, 1 mm. wide; 
calyx tube obconic, 2-3 mm. long, 3-4 mm. 
across, subappressed-pilosulous, subtended 
by two persistent subulate brown subap- 
pressed-pilosulous bracts 1-1.5 mm. long; 
calyx lobes 4, of unequal length, 1 opposite 
pair 3-4 mm. long, 3-4 mm. wide, the other 
pair 2-3 mm. long, 3-4 mm. wide, ovate, 
green, fleshy, persistent, below finely yellow 
glandular-punctate, appressed-puberulous, 
above glabrous, within the tube attached to 
annular disk; petals 4, white, spreading, per- 
sistent, inserted on margin of disk, concave, 
ovate or obovate or elliptic, apex subacute or 
obtuse, base truncate, 6-8 mm. long, 4-5 
mm. broad, membranaceous, minutely glan- 
dular-punctate, ciliate and sparsely puberal- 
