204 
ically isolated subspecies, but not enough 
specimens are known to draw any conclusions 
as yet. 
Alpheus gracilipes Stimpson, I860 
For synonymy, see Banner, 1956. 
localities: Two specimens at E-l-306; 2 
at E-l-387; 3 at E-l-617. 
discussion: Of these 7 specimens, 2 males 
and 2 females had their small chelae intact; 
all 4 had the characteristic fringe of hair over 
the dactylus, but none had their fingers no- 
ticeably broadened. These specimens, there- 
fore, are like the form found in Hawaii and 
not like the form from the Marianas. 
Alpheus paracrinitus var. bengalensis 
Coutiere, 1905 
For synonymy, see Banner, 1956. 
locality: One specimen at E-l-106. 
Edwardsi Group 
Alpheus crassimanus Heller 
Alpheus crassimanus Heller, Reise der . . . 
Novara . . . Zool. Theil 2(8): 107, pi. 10, 
fig. 2, 1865. 
localities: One specimen at E-l-387; 2 
(fragmentary) at E-l-670; 3 at E-2-311. 
discussion: The fragmentary specimens 
were assigned to this species because those 
parts remaining were almost identical to spec- 
imens of similar size from Hawaii. 
One specimen was somewhat unlike the 
rest with the sculpturing on the large chela 
deeper and more abrupt, and the second and 
third legs thinner and more elongate (the 
merus of the third legs is 6 times as long as 
broad instead of the reported 4.5 to 5.0 
times). However similar variation in the large 
chela has been reported for Hawaiian spec- 
imens (Banner, 1953: 134), and for this com- 
parison, the meri of a group of specimens 
PACIFIC SCIENCE, Vol. XI, April, 1957 
collected from a single area in Kaneohe Bay, 
Oahu, Hawaii, were measured, and it was 
found that the length-breadth ratio varied 
from 4.5 to 6.0. 
Alpheus strenuus Dana 
Alpheus strenuus Dana, U. S. Explor. Exped. 
13(1): 543, pi. 34, fig. 4, 1852. 
localities: One specimen at E-2-169; 6 
specimens, Arno, records lost. 
discussion: Some of these specimens, the 
largest I have yet seen, reach a total body 
length of 57 mm.; Coutiere (1905: 913) has 
reported specimens reaching 95 mm., but his 
figures included the chelae. 
The distinction between this species and 
A. audouini described by Coutiere in 1905 
appears to be somewhat questionable. Cou- 
tiere makes the distinction upon the form of 
the anterior carapace and rostrum, and the 
form of the large chela, but the distinctions 
are so fine as to be scarcely apparent in his 
figures. Considering the variations that have 
been reported for other species of the genus, 
these fine distinctions appear to be of dubious 
validity. Probably the best differentiations 
may be found in the antennular base, where 
the median article is 1.5 times the length of 
the distal article in A. audouini and 2 times 
the length in A. strenuus; in the second legs, 
where the first article is 1.65 times the length 
of the second in A. audouini and 1.0 to 1.2 
times the length in A. strenuus; and in the 
merus of the third legs where the length is 
4.3 times the breadth in A. audouini and 3.7 
times the breadth in A. strenuus . Even these 
differences could easily be within the normal 
range of variation for an inherently variable 
species. 
Alpheus pacificus Dana, 1852 
For synonymy, see Banner, 1956. 
locality: Three specimens at E-l-106. 
