Johnstonianidae — Newell 
407 
Grandjean System, with Modifications Terminology Proposed by Various 
Introduced in This Paper Specialists in the Larval Trombiculidae 
solenidioni or Si spur of tarsus I 
solenidion 2 or s 2 . spur of tarsus II 
solenidion 3 or s 3 microfemorala, microgenuala, 
microtibiala (in part) 
solenidion 4 or s 4 microtibiala (in part) 
solenidion of palpal tarsus spur of palpal tarsus 
dorsal eupathid of larval tarsi, or "eupathid at 0.71*7” (etc.) . . subterminala 
distinventral eupathid of larval tarsi, or "eupathid at 0.92 pv” 
(etc.) pretarsala (a misnomer) 
eupathidia of various segments of adult legs (no acceptable equivalent) 
companion seta of dorsal eupathid of larva parasubterminala 
companion seta of Si or s 2 of larva (no equivalent) 
companion seta of s 4 of larva (no equivalent) 
eupathidia of palpal tarsus of larva subterminala of palpal tarsus 
of larva 
eupathidia of palpal tarsus of adult apical setae or apical nude setae 
supracoxal setae (no equivalent) 
vestigial setae microgenuala, microtibiala 
famulusi microspur of tarsus I 
famulus 2 microspur of tarsus II 
bothridia mastifemorala, mastitibiala, 
mastitarsala 
scendants of the "subterminala” of the tarsi 
of the legs are simply called "nude setae” in 
the few cases in which they have been men- 
tioned. "Nude setae” is apparently a catch-all 
category which includes famuli, solenidia, 
eupathidia, vestigial setae, and occasional 
normal setae which lack barbs. What is equally 
remarkable is that setae which are obviously 
morphological equivalents of the larval "sub- 
terminala” show up on segments as far back 
as the telofemur in adults. The term "sub- 
terminala” is therefore totally inappropriate 
to designate these setae, while the name "nude 
setae” is ambiguous. These are all eupathidia, 
and that term can be applied to these setae 
whether they are in larva or adult, on palp or 
leg, on tarsus, tibia, patella or telofemur. The 
"pretarsala” is another eupathid, but it is not 
on the pretarsus, and moreover its ontogenic 
counterparts in the adult are frequently in- 
distinguishable from those of the "subter- 
minala.” The eupathidia are a particular type 
of seta found in a variety of places, and it 
seems unnecessary to apply special names to 
them. 
The "genuala” and "tibiala” are both sole- 
nidia, and nearly all "tibiala” are structurally 
identical with all "genuala.” Why, then, 
should these be given different names, just 
because they are on different segments of the 
legs? Why not also apply separate names to 
the normal setae found on the separate seg- 
ments of the legs? It can also be pointed out 
that certain of the larval "tibiala” have coun- 
terparts which appear on the tarsus of the 
adult. The "spurs” are also solenidia, al- 
though differing structurally from the other 
solenidia of the legs. Giving the various types 
of solenidia different names, confusingly 
parallel in construction in some cases to the 
names of entirely different types of setae 
(femorala, genuala, tibiala, microtibiala, mi- 
crogenuala, microtarsala, subterminala, pre- 
tarsala) complicates the picture unnecessarily. 
What is worse, it obscures the fundamental 
relationships and true differences between the 
setal types. These have been very adequately 
outlined by Grandjean, whose works are to 
be recommended most highly to any who 
have not yet read them. 
During the course of the present studies 
it was found necessary to elaborate somewhat 
upon the solenidial classification in the Para- 
sitengona. The Johnstonianidae, possibly 
more than any other family, show clearly the 
multiplicity of form of these setae, but at the 
