Sphaeniscus Becker and Euphranta Loew of the Oriental and 
Pacific Regions 1 (Tephritidae-Diptera) 
D. Elmo Hardy 2 
Reviews of the genera Sphaeniscus Becker 
and Euphranta Loew of the oriental and Pa- 
cific regions are presented in this study. The 
generic concepts of Euphranta have been re- 
vised and Staurella Bezzi is considered a sub- 
genus of Euphranta . I have placed six species 
in Euphranta ( Euphranta ) and 25 species in 
Euphranta ( Staurella ) . One new species in each 
of these groups is being described from the 
Philippine Islands. The drawings have been 
prepared by Marian S. Adachi, University of 
Hawaii; this help is greatly appreciated. 
Sphaeniscus Becker 
Sphaeniscus Becker, 1908. Berlin Zool. Mus., 
Mitt. 4: 138. 
Spheniscomyia Bezzi, 1913, Ind. Mus., Mem. 
3: 146-147 [Invalid emendation of Sphae- 
niscus Becker]. 
Characterized by having vein R4+5 bare or 
but sparsely bristled; by having four scutellar 
bristles (except for S. neivae (Bezzi) from 
Africa) ; the r-m crossvein situated toward the 
apical portion of cell 1st M 2 ; the arista pu- 
bescent; the cubital cell with a very short 
angular lower apical lobe; the postocular cilia 
and other cephalic bristles all black; the body 
shining black without yellow markings; the 
wings black with a pattern of hyaline spots 
1 Published with the approval of the Director of the 
Hawaii Agricultural Experiment Station as Technical 
Paper No. 320. Manuscript received March 25, 1954. 
2 Entomologist, Hawaii Agricultural Experiment 
Station, College of Agriculture, University of Hawaii. 
including four to five on the hind margin and 
the basal portion of the wing. 
key to species of Sphaeniscus known from 
THE PACIFIC AND ORIENTAL REGIONS* 
1. Wings with four hyaline spots on the 
hind margin (Fig. 1) 2 
Wings with three hyaline spots on the 
hind margin (S. E. Asia) 
quadrincisus (Wied.) 
2. Two hyaline spots in median portion of 
wing (Fig. 1) (Fiji) . .binoculatus (Bezzi) 
No such hyaline spots (Fig. 2) (Pacific 
and Orient) . sexmaculatus atilia (Walker) 
Sphaeniscus binoculatus (Bezzi) 
Fig. 1 
Spheniscomyia hinoculata Bezzi, 1928, Dipt. 
Brachycera Fiji Isl., 115-116. 
This species is closely related to S. sex- 
maculatus. It is distinguished by having two 
round isolated hyaline spots in the middle of 
the wing, one situated at the upper basal 
third of cell 1st M 2 , slightly over vein Mi+ 2 , 
and one at the upper middle of cell R 5 (Fig. 
1) rather than having the distal wedge-shaped 
mark on the hind margin of the wing extend- 
ing through cell R 5 almost to vein R4+5 and 
the wedge-shaped proximal mark extending 
to the middle of cell 1st M 2 (Fig. 2). The 
* Spheniscomyia apicalis Shinji (1939: 322) is appar- 
ently a Trypeta (Ito, 1947). 
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