436 Browne. — A Fourth Contribution to our Knowledge of 
Cone C there were seven distinct meshes arising within the cone, and 
there was also an indication of what might be regarded as a minute and 
deferred mesh, but is a mere local failure of a few elements to become 
tracheides. At this point, between the second and third whorls, the xylem 
is only separated by a single endodermal cell, common to both endo- 
dermes. Five of the meshes are closed, two being of the first and three 
of the second order. At the level of the third whorl one of the strands 
of Cone A passes through the node without giving off a trace. A simi- 
Text-fig. 8. Recon- 
struction of the stele of Cone 
A of E. variegatum. Axial 
xylem black ; traces and 
parenchyma white ; i.a. level 
of insertion of annulus, 
x i 3 t- 
Text-fig. 9. Recon- 
struction of the stele of Cone 
B of E. variegatum. Axial 
xylem black ; traces and 
parenchyma white ; i.a., level 
of insertion of annulus; c.a., 
level at which axial bundles 
cease to present the appear- 
ance of internodal bundles of 
the vegetative axis, x 13^. 
Text-fig. 10. Recon- 
struction of the stele of Cone 
C of E. variegatum. Axial 
xylem black ; traces and 
parenchyma white ; i.a., level 
of insertion of annulufe ; c.a., 
level at which axial bundles, 
cease to present the appear- 
ance of internodal bundles of 
the vegetative axis, x 13^. 
lar irregularity has been observed in E. limosum (Browne ( 1 ), pp. 678-9) 
and in E. maximum (Browne ( 2 ), PI. XIII). 
In E. variegatum , as in the other species studied, certain paren- 
chymatous meshes originating below the cone persist into the latter. These 
will be considered on a later page. 
IV. The Apex of the Cones. 
In Cones A and C of E. sylvaticum the apex of the cone is traversed by 
a single strand, resulting from the fusion of two persistent vascular strands. 
At their junction these two strands enclose a few parenchymatous cells, but 
during the narrowing of the terminal strand, which follows rapidly on its 
constitution, these disappear, and the xylem forms a solid strand, more or 
less circular in transverse section. The structure in these cases is very 
similar to that found in the apices of the cones of E. maximum (Browne ( 2 ), 
p. 242). In Cone B, on the other hand, two strands lying a certain distance 
apart in the parenchymatous ground tissue persist into the ‘ drip-point ’ of 
the cone. The extreme apex of the latter had been injured, and it was not 
