Seedling Structure of Gy mno sperms . //. 207 
phloem and rotation of the xylem, a rearrangement of tissues which was not 
shown by a . Very quickly a junction between the phloem of a" and half 
the bast of a was effected, and in this condition the vascular strands entered 
the axis. A pole of the root-structure was entirely formed by the elements 
derived from the strand a, the other bundle a played no part. This means 
that one bundle (a) is the equivalent of the trace of a whole cotyledon, while 
the other (a) represents the bundle of a subsidiary seed-leaf. 
It only remains to be remarked, that the bundle of the third cotyledon 
first showed the characteristic redistribution of the vascular tissues within 
the hypocotyl, not in the cotyledon. This seedling thus shows three 
cotyledons on the way to become five. 
Series C. This seedling, in many respects, resembled the foregoing 
individual. There were four seed-leaves, the bundles of two of which 
behaved normally ; one as that of a whole-cotyledon, and the other as that 
of a subsidiary seed-leaf, although the xylem of this last strand retained an 
abnormal and isolated position beside one of the poles of the diarch root. 
The two remaining cotyledons fused together at some little distance above 
the cotyledonary node ; so that in transverse section the appearance of this 
single structure resembled the seed-leaf illustrated in Diagram 6. Of these 
two strands one bifurcated very unequally ; that part of the bundle situated 
nearest the other strand being very small (see Fig. 3, PI. XV). The subse- 
quent events resembled those described for Series B sufficiently closely 
to render any further description unnecessary. 
Series D showed similar phenomena, as regards the essential features. 
Pinus sylvestris, L. 
The following table shows the variation in the number of the cotyle- 
dons ; it will be observed that the number varies from 3 to 8, which agrees 
with the observations of Masters 1 ; and that seedlings with six seed-leaves 
are the commonest. Dangeard 2 gives 6-9 as the range in the number 
of the cotyledons, and enters somewhat fully into the structural details. 
No. of 
cotyledons. 
No. of 
seedlings. 
% 
3 
2 
0-21 
4 
28 
2*97 
5 
1 7 1 
l8* II 
6 
500 
52*97 
7 
209 
22-14 
8 
34 
3.60 
Total 944 
The majority of the seedlings examined showed transition-phenomena 
of exactly the same nature as in many of the foregoing plants, especially 
1 loc. cit. 2 Le Botaniste, iii, 1892. 
