252 
Davie . — The Structure and 
directly out from the edges of the palea. Occasionally a gland may be 
found on the adaxial face of the palea, usually about its centre. Curious 
curved hairs are liberally distributed upon the surface of the petiole and 
rachis among the paleae. These hairs are inserted on distended boss-like 
cells of the epidermis. Each hair is made up of eight or ten cells of 
rectangular form. Two of these rise directly up from the boss-like cell, 
while the rest form a tapering series which bends over at its tip almost to 
touch the epidermis. At the top of the curved part two or three glands 
are inserted. These are of the unicellular type found on the paleae. 
Sometimes a gland may be inserted near the tip of the hair, but more 
usually a group is found on the curved portion of the hair (Fig. 9). 
Here and there on rachis, petiole, and on the back of the veinlets of 
the pinnae and pinnules unicellular glands are also found simply inserted 
on the epidermal cells. In the young parts of the plant and on the petiole 
these glands have a hyaline margin, which appears to be filled with a clear 
fluid secreted by the gland (Fig. 10). Right over the top of this secretion 
passes a limiting membrane, which does not, however, appear to be cutinized, 
though it is continuous with the outer layer of the stalk of the gland. In 
the older parts of the plant and on the lower portion of the petiole the 
glands have no hyaline covering, but are limited externally by a thin cell- 
wall, and contain a distinct nucleus and dense cytoplasm (Fig. 11). 
Apparently the secreted fluid disappears from the glands as they grow 
older. The glands closely resemble those found in the tissue of the petiole 
and stock of Nep hr odium filix-mas (de Bary, pp. 89 and 220, and Schacht, 
’63). They give to the plant a very distinct odour, somewhat recalling 
that of Heracleum Sphondylium , and also bearing some resemblance to that 
of Lastrea Oreopteris. In the case of Peranema , however, the glands do 
not show the golden appearance of those in L. Oreopteris , but rather a pearly 
white appearance by reflected light, while they are quite translucent by 
transmitted light. 
On the lower surface of the veinlets in the pinnules are found short 
recurved hairs bearing no glands. They project from the epidermis and 
give the suggestion of little bent pegs hanging down from the pinnules. 
They are multicellular, but their cells are in a single series or row, and are 
some six or seven in number. 
The segmentation of stem and leaf goes on in the usual fashion for 
Leptosporangiate Ferns. The wings of the leaf show the customary marginal 
series of cells (Fig. 12). 
Sorus and Sporangia. 
The mature sorus and sporangium have been described by Mettenius 
(‘ Filices Horti Bot. Lips.’, 1856, No. 40), and by Wallich (‘ Plantae Asiat. 
Rar. ’, p. 42). Drawings of sorus and sporangium after F. Bauer appear in 
