266 
Davie . — The Structure and 
of its vascular anatomy ; the sorus is a mixed one, with no traces of an 
earlier basipetal sequence ; the annulus is vertical in insertion, but slightly 
twisted in its course across the sporangial head ; the spores are markedly 
‘ aspidioid’ ; the prothallus is Cyatheaceous. 
3. Both Ferns show relationships with the members of the genus 
Nephrodium , particularly with Nephrodium filix-mas , Rich., in vascular 
anatomy, the character of glands on the leaves, the sporangial stalks, and 
the spore- markings. 
4. Peranema and Diacalpe are most nearly related to Woodsia and 
Hypoderris , and fall naturally into the Woodsieae-Woodsiinae group of 
Polypodiaceae, which is intermediate between the Cyatheaceae and the 
Aspidieae. 
5. It seems probable that the Aspidieae sprang from a Gradate 
ancestry, and that Peranema and Diacalpe are relatively early members of 
a phyletic drift to the Polypodiaceae, of which the Cyatheaceae are near 
living representatives. 
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