335 
Laboulbenia chaetophora and L. Gyrinidarum . 
bottoms of the pits. In another cell the ‘ fibrils ’ are shown passing around 
the nucleus on their way across. It was thought at first that possibly these 
strands of cytoplasm connected the nuclei of contiguous cells in some way, 
such for instance as Strasburger found to be the case with pairs of nuclei in 
the same cell in Penicillium (Strasburger (’ 02 ), Schiirhoff (’ 07 ) contra ), but 
Fig. 20 lends no support to such a view. It is true, nevertheless, that 
cytoplasmic fibrils can in some instances be traced from the pits to the 
nucleus, and that in degenerating cells they appear to be the most 
persistent parts of the cytoplasm. 
That the nucleus bears a very evident relationship to the organization 
of the surrounding protoplasm at times has been dealt with in some detail 
by the writer in an earlier paper (Faull, ’ 05 ). Hydnobolites and Sordana 
furnish indubitable evidence of this contention. 
Antheridia. 
Careful and repeated search has been made for antheridia, but no 
organs resembling either the exogenous or the endogenous types have 
been discovered in either L. chaetophora or L. Gyrinidarum. So far 
as our knowledge goes these species afford examples of forms that have 
lost their antheridia. Belonging, as they do, to a genus characterized 
by conspicuous flask-shaped simple endogenous antheridia, it is not likely 
that they can have been overlooked, though that is not outside the range 
of possibilities. The cytology of both types of antheridia has been studied 
in other forms, and an account of the phenomena observed will be very 
shortly published. 
Procarp. 
The procarp has its origin as a uninucleate terminal cell of a lateral 
branch of the receptacle (PL XXXVIII, Fig. 2i). The two basal cells of the 
branch represented in Fig. 21 are daughters of the obliquely septated ‘pri- 
mordial cell of the perithecium and give rise to the perithecium proper. 
The course of development of the perithecium has been very care- 
fully worked out by Thaxter, and so need not be recapitulated in detail 
here. It may be stated, however, that transverse sections wholly verify 
his statements that the number of rows in both the outer and inner walls 
is four, and that the rows of the two walls alternate with one another 
(Figs. 33-6). The inner is of later origin than the outer ; in fact, does 
not appear until the latter is nearly complete and the procarp fully 
formed. Both are clearly outgrowths of the basal cells of the perithe- 
cium. They have been repeatedly seen in longitudinal sections in all 
stages of development, and their protoplasmic connexions with the basal 
cells demonstrated (Fig. 25). 
