336 Fault . — The Cytology of 
The mature procarp consists of three parts, the carpogonium, the 
trichophoric cell, and the trichogyne. The trichogyne is very highly 
developed in L. chaetophora and L. Gyrinidarmn. It is an elaborately 
branched, frequently septated structure consisting of uninucleate cells 
(Figs. 30, 31). It, too, has been figured and described by Thaxter, and 
little further can be added. 
The possible functions of trichogynes that appear no longer to serve 
as receptive organs of spermatia have been recently fully discussed 
by Vuillemin (’ 07 ) and Brooks (TO), but so far as I know no experi- 
mental evidence has been adduced in confirmation of any of them. 
The sequence of cell-formation in the carpogenic cell is strictly 
comparable to that described by Thaxter for L. elongata. The tricho- 
phoric cell undergoes no further division, though its nucleus as described 
below may play a very significant role. The carpogenic cell divides into 
three superposed cells, the superior and inferior supporting cells (^..r. and 
i.s.) and the ascogonium (am.) which lies between them. The ascogonium 
may again divide transversely, the lower half constituting a secondary 
supporting cell, while the upper usually divides by an oblique partition into 
the ascogenic cells (asc.) from which the asei (as.) bud out. In some plants 
this division does not take place, so that there may be but one ascogenic 
cell. Two, however, is the characteristic number, and never more than two 
have been found in the species here described. Two vertical rows of asci 
now bud out from each of the ascogenic cells. An account of their further 
development will be given in a subsequent section, and the cytology of the 
procarp only dealt with here. 
The carpogenic cell is at first uninucleate, a relatively large nucleus 
with a large nucleolus occupying the centre of the cell (Fig. 22). The 
trichophoric cell is also uninucleate (Fig. 22). The single, centrally placed 
nucleus of the carpogenic cell now apparently divides ; at all events it is 
succeeded by a pair of smaller nuclei (Fig. 24). I have not observed its 
actual division, but the nucleus represented in Fig, 23 appears to be in an 
early prophase stage. The nucleus of the trichophoric cell very soon moves 
down next to the carpogonium, as shown in Fig. 25, and undergoes a homo- 
typic mitosis. The central bodies of the spindle are large and disc-like, but 
it is difficult to make sure of the number of chromosomes. At this stage, or 
even earlier, the partition separating the two cells disappears. Thus in 
Fig. 26 there is the appearance of a long single cell in which lie four nuclei, 
A, B, c, D, enumerating from the bottom up. A and B presumably are 
daughters of the carpogenic nucleus, C and D of the trichophoric. The 
lower end of this cell with nucleus A is eventually cut off as the inferior 
supporting cell, and the upper end with D as a ‘ restored ’ trichophoric cell. 
But it was impossible to determine the exact sequence of septa-formation. 
The middle pair, B and C, divide after a short period of growth, and 
