446 
Stiles . — The Podocarpeae. 
with 1 6, Saxegothaea and Microcachrys each monotypic, and Pherosphaera 
with 2 species. 1 In all genera and species a single ovule is borne in a 
median position on the megasporophylh The megasporophylls may be 
closely aggregated in well-defined cones as in Saxegothaea and Micro- 
cachrys, the cone may be a reduced one as in Pherosphaera , a still more 
reduced one as in Phyllocladus, or the sporophylls maybe several in number 
as in some species of Dacrydium and Podocarpus ; lastly, they may occur 
singly or in pairs as in other species of these two genera. Again, the ovules 
may be pedicillate as in Podocarpus , or sessile on the sporophyll as in the 
other genera. In this case, they may be completely reversed as in Saxe- 
gothaea and Microcachrys , only partially reversed as in some species of 
Dacrydium , or erect as in Phyllocladus and Pherosphaera. Finally, they 
may have only one integument (. Pherosphaera ), two integuments with the 
outer only partially surrounding the inner, or the outer integument may be 
symmetrical (. Phyllocladus ). In spite of these differences there can now be 
no doubt that the forms here included form a natural family. Other evidence 
supporting this conclusion will be given in the course of the paper, and 
indeed such a view is now almost universally adopted. 
The following genera and species have been examined : 2 
1. Saxegothaea , Lindl. 
The single species 5 . conspicua y Lindl., has been described as 
Squamotaxus Albertiana , Senilis, but it would appear that no one but the 
author of the name has ever employed it. The plant is a small tree of 
Yew-like habit, native of the Chilian Andes, and is monoecious. A case 
has, however, been recorded in which some of the branches on a tree were 
mostly microsporangiate and others ovulate. 3 
2. Microcachrys , Hook. f. 
The only species, M. tetragona , Hook, f., was for long confused with 
Athrotaxis , Dacrydium , and Pherosphaera . The male cones were first 
described by Sir W. J. Hooker in 1843 as A. tetragonal and again in 
1845 by Sir J. D. Hooker as Microcachrys tetragonal the female cones 
then described under that name being those of Pherosphaera . Both 
male and female cones were described by Archer in 1850 as Pherosphaera . 6 
1 The New Caledonian genus Acmopyle (Pilger, ’03, p. 117), with its single species^. Pancheri , 
probably belongs to this order, but so little is known of it that it seemed doubtful whether any 
useful purpose would be served by introducing it into this paper. 
2 The nomenclature of Pilger’s monograph (’03) has been followed throughout as the most 
recent and most complete systematic account of the Podocarpeae. Further information on external 
features of the species is to be found in that work. Photographs showing the habit of some species 
of Podocarpus and Dacrydium are given by Laing and Blackwell (’06). For photographs of 
Dacrydium Franklini , Pherosphaera Fitzgeraldi , Phyllocladus rhomb oidali s , Podocarpus elatus , and 
P. spinulosus , see Baker and Smith (TO). 
3 Noren (’08), p. 102. 4 Hooker, Sir W. J. (’43), Tab. 560. 
5 Hooker, Sir J. D. ('45), p. 149. 6 Archer (’50), p. 52* 
