Stiles. — The Podoccirpeae. 451 
the ring of bundles comprising the stele. Probably the condition of 
affairs is essentially the same as in some of the Araucarieae. 1 
Podocai'pus amarns is essentially the same as P. andinus as regards 
the structure of the young stem. This is, as usual, a ring of bundles with 
a single resin canal outside each bundle. The case is, however, a little 
complicated as each leaf bundle is accompanied by three resin canals 
(Text-fig. 1, c ). These three canals run down into the stem with their 
corresponding bundle, but the two lateral ones ultimately fuse with neigh- 
bouring canals, so that the bundle is left with a solitary canal. In this 
stem stone cells are extremely abundant in the pith, though before secondary 
growth had commenced only a few solitary ones were observed in the cortex. 
As in P. nagi , the primary xylem is practically limited to the protoxylem ; 
PI. XLVII, Fig. 1 7, shows a primary bundle in the stem just as the 
interfascicular cambium is forming. 
In the stem of Dacrydium cupressinum there is a layer of sclerenchyma 
outside and in contact with the phloem. In the stems examined in the 
course of this work the sclerenchyma was two or three layers thick. The 
primary medullary rays in this stem are narrow, in some cases being only 
two cells wide. A ring of resin canals occurs in the cortex, as in the species 
of Podocarpus examined. 
Dacrydium Franklini is the only other species of the genus examined. 
There is the usual ring of vascular bundles, but unlike D. cupressinum , and 
although resin ducts are present in the leaves, canals are absent from the 
stem (Text-fig. 1, e). The same is apparently the case in Microcachrys. 
B. Secondary Xylem. 
The accounts we possess of the structure of the stem in the Podocarpeae 
are mostly confined to accounts of the secondary wood. They are those of 
Bertrand 2 on Podocarpus and Dacrydium (treated as a section of Podocarpus ) , 
and of Penhallow 3 on Podocarpus macrophyllus . Beust 4 examined Saxe- 
gothaea , and Gothan 5 Saxegotkaea, Dacrydium spp., and Podocarpus spp. 
Baker and Smith have recently published photographs of the wood of 
Dacrydium , Pliyllocladus , and Podocarpus . 6 The writer has previously 
examined the stem of Saxegotkaea , 7 and in conjunction with Mr. Brooks 
that of Podocarpus spinulosus . 8 Miss Gerry 9 has recorded some observations 
on the occurrence of ‘ bars of Sanio ’ in these plants. 
Bertrand’s observations are largely in terms of Taxus . He found that 
in Podocarpus wood parenchyma was often present in greater quantity than 
1 Seward and Ford (’06), p. 351. 2 Bertrand (’74). 
3 Penhallow (’04) ; (’07), p. 216. 4 Beust (’84), p. 38. 
6 Gothan (’05), p. 57. 6 Baker and Smith (’10), pp. 405, 428, 439. 
7 Stiles (’08), p. 210. 8 Brooks and Stiles (’10), p. 306. 
9 Gerry (’10), p. 119. 
