457 
Stiles. — The Podocarpeae . 
D. Franklini quite confirm these statements. The juvenile form of leaf of 
D. cupressinum is tetragonal in transverse section, with a sclerenchymatous 
hypoderm continuous round the leaf, except in the region of the stomata. 
These occur in two bands towards those corners of the tetragonal leaf which 
are morphologically the margins. Palisade tissue is continuous all round 
the leaf, there being perhaps a slight development of spongy tissue in the 
corners corresponding to the margins. 
There is a single vascular bundle with a well-marked resin canal. 
A well-marked sheath of parenchymatous cells surrounds the bundle and 
resin canals, a single layer of similar cells separating the phloem and the 
canal. Well-marked transfusion tissue is absent, but some lignified paren- 
chymatous cells about the xylem probably represent it. These cells, 
although in the position of centripetal xylem, have exactly the appearance 
of parenchymatous cells, except for their lignified walls, and in this case 
certainly do not appear to be modified xylem-elements (PL XLVII, Fig. 19). 
The leaf of the adult form is more flattened than the juvenile form, and is not 
markedly tetragonal in section (Text-fig. 2 ,e). The structure, otherwise, 
is almost identical with that of the juvenile form. The stomata are in 
a similar position, but more numerous, while the lignified cells probably 
equivalent to transfusion cells occur in two rather lateral groups (PI. XLVII, 
Fig. 20). 
5. The leaf of Microcachrys tetragona has been described by Thomson, 1 
with whose account my own observations agree. There is a distinct 
palisade with sclerotic hypoderm on the lower surface, while hypoderm on 
the upper surface is wanting. The stomata, as in Dacrydium, occur on the 
upper surface only. The single resin canal is wholly foliar, not being 
carried down into the stem. 
6. Phyllocladus. The writer has not had an opportunity of examining 
leaves of this genus. Mrs. Arber 2 has described those of Ph . alpinus , 
Ph. tricho memo ides, and Ph. rliomboidalis . They are, as would be expected 
in such a reduced leaf, of very simple type. On the whole they most nearly 
recall those of Microcachrys , except that no hypoderm or palisade is present. 
A single resin canal is present under each bundle. There is no differentiation 
of tissues in the mesophyll of the leaf. Mrs. Arber did not observe stomata 
in Ph. alpinus and Ph. trichomanoides , but a few were present in Ph. rhotn- 
boidalis. 
7. Nageia. Of the species of the Nageia section the only one whose 
leaves have been examined in the course of this investigation is Podocarpus 
nagi. In this species no stomata were observed on the upper surface of the 
leaf, a character in which, according to Bertrand, it differs from other species 
of the section. In the leaves from Tokyo examined there is a scleren- 
1 Thomson (’09 2 ), p. 349. 
2 Robertson (’06), p. 260. 
