460 
Stiles . — The Podocarpeae. 
of one leaf may vary from one to four. 1 The stro'bili of Phyllocladus are 
borne in clusters of 2 to 8 in the axils of scales at the tips of the lateral 
branches. 2 In Microcachrys , on the other hand, the male cones are terminal 
on an ordinary leafy shoot. 3 
Coulter and Chamberlain 4 have remarked on the relation of leaves 
and strobili in the Taxaceae as being such, that if the axis were shortened, 
a compound strobilus would result. Although this statement does not 
hold generally for the Podocarpeae (as witness the case of Microcachrys ), yet 
it is nevertheless true for Saxegothaea and all the species of Podocarpus 
examined by the writer. The resemblance to a raceme of the cone-bearing 
branch previously noticed in the case of Saxegothaea 5 is even still more 
noticeable in the case of Podocarpus andinus (PI. XLVI, Fig. 5). 
In Microcachrys , where the foliage leaves are arranged decussately, the 
sporophylls are arranged in alternating whorls of four. In all other genera, 
as far as they have been examined, the sporophylls are spirally disposed on 
the cone axis. At the base of the cone occur a few small spirally arranged 
bracts. 
The writer has only had an opportunity of examining the male cones 
of Saxegothaea , Microcachrys , Podocarpus andinus , P.polystachyus , P. spinu- 
losus , and P. latifolius , but descriptions are also available of a few other 
species. These cases are, however, sufficient to show that a good deal of 
variation in shape and mode of attachment of sporangia exists in the order 
(Text-fig. 3). 
In P. andinus each sporophyll is attached to the axis of the cone 
by a short narrow stalk, which terminates in the relatively large triangular 
upward-projecting portion ending in a broad point. The two sporangia are 
inserted on this on the side near the cone axis, one on each side of the 
stalk and partly attached to it, and are not in contact with one another 
(PI. XLVI, Fig. 6, and Text-fig. 3, a). In Podocarpus latifolius the extent 
of fusion is greater and the sporangia are radially elongated ; a tangential sec- 
tion of the cone shows that they are fused together for the upper half of 
their vertical width (Text-fig. 3, b). In Saxegothaea , and presumably also 
in Podocarpus Totara Hallii , 6 the two sporangia are fused together for the 
greater part of their vertical width (Text-fig. 3, c). The same is the case 
in Microcachrys , but here a large cavity is present between the two 
sporangia and below the phloem of the vascular bundle (Text-fig. 3, d). 
In Saxegothaea the line of dehiscence of the sporangium is transverse, 7 and 
transverse dehiscence of the sporangium of Dacrydium cupressinum is 
figured by Richard and by Lambert, 8 and described for D. elatum and 
1 Brooks and Stiles (’10), p. 307. 2 Coulter and Chamberlain (’10), p. 319. 
3 Thomson (’09 1 ), p. 26, PI. I, PI. II, Figs. 1 and 2. 
4 Coulter and Chamberlain, 1. c. 6 Lindley (’51). 
6 Burlingame (’08), PI. VIII, Fig. 6. 7 Stiles (’08), p. 212. 
8 Richard (’26), PI. II, Dacrydium cupressinum , Fig. K; Lambert (’32), Tab. 69. 
