470 
Stiles . — The Podocarpeae. 
VI. Podocarpus. 
i. Eupodocarpus} The early development of the female fructification 
has been traced by Strasburger 2 in the case of Podocarpus chinensis , 
Wall. (P. macrophyllus , subsp. maki , Sieb.). The ovulate structure is 
described as arising in the axil of the sporophyll, the integument as 
growing up round the nucellus, while the horseshoe-shaped aril arises 
later. 
Some young fructifications of P. latifolius collected by Mr. Saxton 
yielded the youngest stages of my material. The ovular stalk is short and 
the ovule at its apex is almost perpendicular to the megasporophyll, and is 
not reversed. The integument and epimatium are only free at the tip 
(PI. XLV'I, Fig. 1 1, and Text-fig. 8,^-). In this species the ovular stalk never 
elongates very much ; by the time the reduction division takes place the 
ovule is, however, completely inverted (Text-fig. 8, //, k , and cf. with /). 
Later the bases of the four ‘ cone ’ scales become fused together into one 
fleshy mass. 
The ovulate structure of Podocarpus macrophyllus is like that of P. 
latifolius in general appearance, but two fertile megasporophylls are perhaps 
of more frequent occurrence than in that species. The fleshy receptacle is 
formed in this species as in the last (PL XLVI, Fig. 10). The seed-coat is 
differentiated into two layers ; a harder and thicker outer layer which, 
however, cannot be described as woody, and a softer inner layer which 
cannot be aptly described as either fleshy or membranous. 
The vascular structure of the female fructification has already been 
described by Mr. Brooks and the writer 3 in the case of Podocarpus spinulosus, 
(Sm.) R. Br., P. alpinus , R. Br., and a third species. These observations 
have now been supplemented by further ones on P. macrophyllus and 
P. latifolius . 
In P. macrophylhis , as in species previously examined, a single bundle 
supplies the megasporophyll, while the ovular supply below the point at 
which the stalk becomes free from the sporophyll consists of two vascular 
bundles, the xylem and phloem of which are inversely orientated as com- 
pared with the xylem and phloem of the sporophyll bundle. These two 
bundles, by division of one or both, give rise to a row of three or four small 
bundles (Text-fig. 4, a-d ). They are accompanied by a row of small 
canals, about one canal to each bundle, though this relation is not strictly 
maintained. Towards the chalazal end of the ovule, if the supply bundles 
number four, one of the bundles at the end of the row may die out, or bend 
round the ovule in a tangential direction and descend towards the micropyle. 
1 For figures of the female fructifications of Podocarpus chilinus, P. elongatus, and P. coriaceus, 
see Richard (’26), PI. I ; and for photographs of these structures in P. elatus and P, spinulosus , see 
Baker and Smith (’10), pp. 432, 444. 
2 Strasburger (79), p. 77 . 
3 Brooks and Stiles (’10), p. 311. 
