5 1 8 Gibbs. — On the Development of the 
differentiation of the megaspore and the first free cell-divisions in the 
enlarging prothallus. 
Pre-fertilization shows the female gametophyte filled with endosperm 
and archegonial initials, or fully developed arch ego nia at the apex of the 
prothallus. In pre-fertilization the pollen-tubes have penetrated the 
stigmatic apex of the nucellus, and with the male prothallus rest over 
the apex of the female gametophyte (Fig. 18, PI. L). 
In the following pages, the $ inflorescence of Podocarpus is described 
as a fully organized strobilus, thus following Bennett and Brown (2), 
Sperk (50), Bertrand (3), and Tison (57). As this point of view has not 
previously been accepted by the generality of botanists, it is perhaps as 
well to give a key to homologize familiar terminology in this genus, as given 
by Pilger in his Monograph, with that used in this paper. 
The different parts of the female flower and axis have, up to the 
present, always been looked upon as so distinct that even portions of the 
same organ exact different names. 
Inflorescence. Female 
flower. 
Branch in D aery carpus, 
Nageia,and. Stachy carpus. 
Scales . 
Scale leaves 
Receptaculum . 
Carpel . 
j Epimatium 
\ ‘ Stumpfer Fortsatz ’ 
Strobilus. 
Stalk in Eupodocarpus. 
Peduncle. 
Scale leaves of the peduncle in Dacrycarpus , 
Nageia , and Stachycarpus. 
Bracts. 
Swollen bract bases in Dacrycarpus and Eu- 
podocarpus. 
Lamina of fertile bract. 
Lamina of the ovuliferous scale. 
Conical apex of the ovuliferous scale ( = semi- 
circular ridge of P. vitiensis ). 
By the fertile bract is meant the bract of the strobilus in the axil 
of which the ovuliferous scale arises. 
It must be explained that in Pilger’s terminology ‘ receptaculum ’ 
stands for the bases of all the bracts of the cone when subsequent swelling sets 
in, as is the case in Eupodocarpus and to a lesser degree in Dacrycarpus . 
When this secondary swelling does not take place as in S tacky - 
carpus (PI. L, Figs. 28 and 29), though the bract bases are equally well 
developed in the young stages (PI. L, Figs. 25 and 27), in the maturer 
stages the laminae are caducous, instead of persisting as points as in 
the two sections above quoted ; the bract bases are therefore spoken of as 
‘ scars ’. 
The ‘ carpel * is described by Pilger as developing out of one of the 
scale leaves. That is to say, one portion only, i. e. the lamina, is spe- 
