532 
Gibbs . — On the Development of the 
which completely covers them, and are borne on peduncles about 5 mm. 
long, which may either terminate a shoot, as Tison figures for Saxegothaea 
( 57 , PL IX, Fig. 6), or they may form little lateral branches, as in P. imbri- 
cata. The peduncle is clothed with imbricating scale leaves (Fig. 10, 
/.), of which those immediately subtending the strobilus are larger and 
more spreading (Figs. 10 and 11, s. /.). 
The bracts may be two (Fig. 11, brs.) or three (Figs. 10 and 12), 
with long terete free laminae, which remain erect or open out. The 
swollen bases are verruculose as in P. imbricata. There is one fertile 
bract, the lamina of which is fused with the ovuliferous scale to the point 
of insertion of the ovule (Figs. 9, 10, and 13). The extreme apex of the 
lamina is more or less free in the younger stages (Figs. 9 and 10), but 
becomes completely fused as growth continues (Figs. 12 and 13). Fig. 10 
is in the pollination stage. 
Histology. A longitudinal section of the ovule shows the lamina of 
the bract with its normally orientated vascular bundle ending in a group of 
tracheides (Fig. 9, v. b. br). The epidermis and hypodermal sclerotic layer 
is continuous with that of the lamina of the ovuliferous scale as in P. imbri- 
cata, The hypodermal layer, as in that species, also characterizes the 
foliage leaves, where it is only interrupted under the stomata which occur in 
four lines, two on each surface of the leaf. This layer is recorded by 
Van Tieghem ( 59 ) for the leaves of both these species. The inverted 
vascular supply of the ovule runs down the ovuliferous scale in two bundles, 
as previously described, forming a ring round the base of the nucellus 
(Figs. 9 and 13), but in P. dacrydioides they were not seen to send branches 
into the lamina of the scale. These bundles run separately, as in P. imbri- 
cata, to the base of the bracts, gradually rotating as they descend, till they 
insert themselves normally on to the ring of bundles of the strobilus. They 
are each accompanied by a resin canal with epithelium. These canals, 
in the region of the ovule, limit the tissue of the integument from that 
of the ovuliferous scale (Figs. 9 and 13, r. c.). 
Pilger ( 40 , p. 23) describes the upper portion of the integument as 
being very thin in this species and the large resin canals as lying in the 
outer seed-coat (‘ aussere Samenschicht ’), presumably the ‘ epimatium 
i. e. lamina of the ovuliferous scale. In the younger stages (Figs. 9 and 13, 
r. c.) it will be seen that there is nothing abnormal in the texture or size of 
the integument or the position of the resin canals, which are, as Pilger 
states, very large in this species. 
The integument shows the usual differentiation of cells, those in the 
region of the base being marked by starch and tannin contents which also 
characterize the epidermis of the free apex at the micropyle (PI. LI 1 1 , 
Figs. 77 and 78). 
The nutritive zone of tannin cells is present at the base of the nucellus, 
