537 
Female Strobilus in Podocarpus. 
the strands. These resin canals must branch independently, as more canals 
appear in the lamina surrounding the ovule than there are vascular strands 
(PI. LIII, Fig. 73). 
In longitudinal section of the pre-fertilization stage the branches from 
the inverted vascular system surround the ovule in the lamina of the scale 
(PI. L, Fig. 17 , v. b. 0. s.) and are seen running down to the micropyle. These 
strands end in the usual group of tracheides (Fig. 17, tra.). This branching of 
the vascular system to serve the lamina of the scale is also present in P. im- 
bricata (PI. XLIX, Fig. 6 , v. b. o. s.). Several strands of phloem elements, of 
narrow elongated cells with densely staining contents, pass from the phloem 
of the bundles across each zone of nutritive tannin cells to the base of the 
nucellus (PL L, Fig. 17, ph . str.) as previously described in transverse section 
(PI. LIII, Fig. 73), where they show on both sides penetrating the zone of 
tannin cells of the integument. The strand rising a little above the bract, 
which runs parallel to the vascular bundle for a short distance (PI. L, 
Fig. 1 7), was present in every ovule sectioned. A phloem strand penetrating 
to the nucellus is also present in the ovules of Agatkis , connecting them with 
the vascular supply, a fact which has been recorded by Worsdell. 
The usual zone of starch and tannin cells limits the base of the integu- 
ment , of which the apex shows elongated spirally thickened and pitted cells 
(Fig. 17, tig. c. inti). 
The base of the nucellus is marked by another zone of tannin cells 
(Fig. 17, tan. and st.c.). The nucellus has a markedly long beak formed of 
serial rows of cells (Fig. 18, nuc .) which are not thickened but merely 
crushed at the extreme apex (Fig. 18, stig. api). This beak is penetrated 
by pollen-tubes which run more or less vertically down it. In one case the 
cA prothallus was seen in the archegonial cavity (Fig. 18, cd pro .), containing 
two generative nuclei with stalk and tube nucleus. In this stage, according 
to Lawson in Sciadopitys verticillata ( 30 ), there is an interval before actual 
penetration into the archegonia. 
The nucellus is limited on the interior by two or three tapetal layers 
with disorganized cells surrounding the gametophyte (Fig. 17, tap.). The 
$ gametophyte has absorbed about two-thirds of the nucellus and (Fig. 17, 
pro.) shows a prolongation into the beak of the latter. 
In all material obtained, though much contracted, a megaspore mem- 
brane is traceable (Fig. 17, m. m.). 
§ Stachycarpus. 
Receptaculum nullum ; flores feminei spiciformes vel ovula 1-2 ad apicem 
ramulorum abbreviatorum. 
P. spicata. 
Morphology. P. spicata , the Matai or Black Pine of New Zealand, 
is a very graceful tree, with a round and densely branching crown, and 
