Female Strobilus in Podocarpus. 
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EXPLANATION OF PLATES XLIX-LIII. 
Strob ., strobilus; ped., peduncle ; s., scale ; bud s., bud scales ; leaf ; s.l ., scale leaves ; br., bract ; 
s.br., sterile bract; f.br ., fertile bract; b. br ., base bract; lam ., lamina; ap. bud, apical bud; tip., 
apex; 0. s., ovuliferous scale ; mic., micropyle ; i., integument; nuc., nucellus ; 0., ovule; v.b.f.br., 
vascular bundle, fertile bract ; v. b. o.s ., vascular bundle, ovuliferous scale ; x., xylem ; ph., phloem ; 
ph. sir., phloem strands ; #a,tracheides; r. c ., resin canal ; mes., mesophyll ; c ., cells ; stone c ., stone 
cells ; st., starch ; t., tannin ; h.f., hypodermal fibres ; e., epidermis ; stom., stomata ; idio., idioblasts ; 
tis., tissue; lig., lignified ; m.m., megaspore membrane; pro., prothallus ; meg., megaspore; tap., 
tapetum; arch., archegonia ; in., initials; n., nucleus; e. n., egg nucleus; j.c., jacket cells; n.c., 
neck cells; p.g., pollen-grain; p.t., pollen-tube; nuc. cap , nucellus cap; pro. emb ., proembryos; 
sus., suspensor ; cm. cyl., central cylinder; dig. c., digested cells; cots. , cotyledons; pi., plerome; 
plu ., plumule ; hyp., hypocotyl ; rt. cap, root-cap ; pro. cam. str., procambium strands ; cor., cortex ; 
stig. ap., stigmatic apex; gen. n., generative nucleus ; stalk n., stalk nucleus ; tube n„ tube nucleus ; 
cup. /., cupressoid; imb ., imbricate; epi., epithelium. 
PLATE XL 1 X. 
Fig. 1. P. imbricata. Branch showing strobili in two stages : this year’s with unmodified 
bracts, in early pollination stage ; last year’s with bract bases swollen at the expense of the lamina, 
in pre-fertilization stage. Nat. size. 
Fig. 2. Strobilus with two bracts, each fertile; bract bases swollen and verruculose; scale 
leaves subtending strobilus removed, x 16, red. \. 
Fig. 3. Strobilus with two bracts, one fertile; showing the apex of the fertile bract unusually 
free and prolonged, and larger size of the scale leaves of the peduncle immediately subtending the 
strobilus. x 16, red. f. 
Fig. 4. Strobilus with four bracts, one fertile, with apex still free, in fertilization stage ; the 
peduncle is covered with imbricating scale leaves, those subtending the strobilus showing less 
modification as the latter increases in size, x 8, red. |. 
Fig. 5. Sterile strobilus, showing base of ovuliferous scale swollen and verruculose (Singa- 
pore). x 8. 
Fig. 6. Longitudinal section through ovuliferous scale and ovule in pre-fertilization stage, 
showing lamina of fertile bract, with normal vascular bundle, fused with the lamina of the scale, with 
inverted vascular system, both enclosing the ovule ; epidermis, hypodermal layer of fibres, and 
mesophyll with lignified idioblasts are continuous for both ; the integumental tissue is limited by 
the resin canals and shows a zone of nutritive cells, i.e. tannin and starch; lignified cells occur in the 
apex, at the micropyle ; the nucellus shows also a zone of nutritive cells, and starch is deposited 
in the cells of the apex ; a tapetum showing continual breaking down of tissue limits the nucellus on 
the inside ; the female gametophyte is invested by a two-layered megaspore membrane, and shows 
archegonia ; the cells of the prothallus are binucleate. x 16, red. |. 
Fig. 6 a. Archegonium with contracted protoplasm and resting egg nucleus, showing neck 
cells at apex, surrounded by jacket cells, x 625. 
Fig. 6 b. Two archegonia in one jacket, seen in a prothallus of another species, x 625. 
Fig. 6 c. Idioblasts in mesophyll tissue. 
Fig. 7. Strobilus with two bracts, one fertile, from the subsummit zone of Kinabalu (11,500 ft.), 
in which the scale leaves do not spread out below the bracts, as in Fijian and Buitenzorg material; 
pollination stage ; the apex of the lamina of the fertile bract projects above the ovuliferous scale, 
x 16, red. j. 
Fig. 8. Longitudinal section of strobilus in pollination stage or after ; the fertile bract shows 
distinct apex, which becomes obscured as ovule matures ; the nucellus is contracted, but pollen-tubes 
can be traced in the apex, x 16, red. |. 
Fig. 8 a. Hypodermal layer of fibres in the lamina of the fertile bract and that of the ovuliferous 
scale, with epidermis, x 6 25. 
Fig. 8 b. Verruculose outgrowths on the swollen bract bases, the epidermis showing tannin 
contents ; no hypodermal layer of fibres, x 625. 
P p 2 
