27 
Anatomy of the Ophioglossaceae. 
positions of these organs is also seen in the accompanying outline figure 
(Text-fig. 6 , f). While, as stated in my earlier paper, the relative position 
of the primary organs of the embryo appears to be the same as is described 
for the embryo of Botrychium virginianum , an essential difference comes out 
when all the sections of the series are considered, and especially when the 
embryo is reconstructed. A number of sections of the series are represented 
in Text-fig. 6. The plane of section had passed at right angles to the 
insertion of the suspensor, which was somewhat flattened by the expansion 
of the embryo. The suspensor, probably the lower tier, is seen in Text-fig. 
6, A and B at j. From it the embryo widened out suddenly, and the other 
sections figured allow of the foot (f), root (r), and shoot ( sk ) being followed 
through the series. The apex of the root is in the section D, while 
Text-fig. 6. Outline figures of a number of longitudinal sections of the upper portion of a 
prothallus bearing a mature embryo. The prothallus, as usual, bore antheridia (an) as well as 
archegonia (ar). The embryo is cut in the longitudinal plane of the erect shoot and the figures 
commence at the side where the suspensor is attached, s, suspensor ; f foot ; r, root ; sk, shoot ; 
cot, first leaf; si, stem. The central vascular system is indicated in D, E, and F. 
the cotyledon (cot) and the apex of the stem (st) are distinguishable in F, 
which is median for the upwardly directed shoot. The position of the 
vascular strand in the shoot and root can be gathered from the sections in 
D, E, and F. 
The reconstructed embryo seen from the suspensor side is represented 
in Text-fig. 3, C. It shows the suspensor (s) and the result of development 
of the embryo proper. The great difference from the smaller embryos 
described is that the shoot with further growth has become vertical. The 
position of the first leaf (cot) and of the stem-apex (st) are indicated, as well 
as the development of hairs over the apical region. The root (r) has grown 
downwards and towards the side away from the mass of the prothallus, 
while the large foot (/) has extended into the latter. Comparison of the 
three embryos in Text-fig. 3 will show what a considerable readjustment 
