T vigo nocarpus Skorensis , sp. nov . 55 
significance that in neither species has the micellar system been traced 
beyond the plane of insertion of the pollen chamber. 
(b) The Sarcotestal System. 
In several sections of the University College series S. 31 #-S. 31, and 
perhaps in section R. 1161, h of the Manchester series, bundles in various 
conditions of preservation are to be found at the extreme periphery of the 
sarcotesta proper, where the outermost lacunar tissue abuts upon the 
limiting layers containing the radial sclerotic plates. The only parallel 
amongst nearly allied seeds to so external a position is to be found in 
Pachytesta } where, however, if we regard the exotesta as representing 
sarcotesta and sclerotesta combined, 2 the bundles are only far out if 
considered in relation to the testa as a whole. 
Owing to the incomplete preservation of the periphery, it is hardly 
surprising that in no slide are more than three sarcotestal bundles present at 
one and the same time. In several of the University College series, S. 31, a- 
S. 31, £, two bundles are cut in successive sections, and both of these are 
fortunately situated between the same pair of major ribs, and also occupy 
the same positions relative to them. Altogether, four distinct bundles can 
be distinguished in this series, and two possible bundles are also recognizable 
in one of the Manchester sections (R. 1161,/?). In every case these sarco- 
testal vascular strands are situated in the radial plane midway between 
a major and a minor rib, and though the bundles are not present, in any one 
of our specimens, on all three faces, yet the occurrence of two, and two only, 
in several successive sections between the one pair of major ribs seems 
to point conclusively to the total number as being six, symmetrically 
disposed around the circumference of the seed. 
The sarcotestal system was then identical as to the number of its com- 
ponents, and almost so as to their arrangement, with that which has been 
described for Trigonocarpus Parkinsoni and Polylophospermump only 
differing in the latter respect with regard to the proximity of the bundles to 
the sclerotic shell. 
The structure of an individual strand is shown in S. 31, i (PI. V, Fig. 17), 
where, owing to the curvature of the surface, the section which is oblique to 
the axis of the seed passes transversely through a sarcotestal bundle 
exhibiting extraordinarily perfect preservation. As can be seen from the 
figure, the bundle is slightly more extended in the tangential than in the 
radial direction (0*39 mm. x 0*17 mm.) and consists of some seventy xylem 
elements, of which the smallest occupy the central region (PI. V, Fig. 17, 
pt. x). The latter, which probably represent the protoxylem, comprise some 
1 Renault, loc. cit. 
2 Oliver : On Gymnospermous Seeds, New Phytologist, vol. i, No. 7, 1902, p. 148. 
3 Oliver, Ann. Bot., vol. xxi, No. Ixxxii, 1907, pp. 303-4. 
