On Medullosa centrofilis, a New Species of Medullosa 
from the Lower Coal Measures. 
BY 
E. de FRAINE, D.Sc., F.L.S., 
Assistant Lecturer in Botany , University of Wales, Aberystwyth. 
With Plate XV and five Figures in the Text. 
Contents. 
I. 
Introduction . 
page 
. 251 
VII. 
Affinities .... 
PAGE 
. 261 
II. 
General Structure 
. 252 
VIII. 
Diagnosis .... 
. 263 
III. 
The Steles 
• 254 
IX. 
Summary .... 
. 263 
IV. 
The Leaf-traces 
. 258 
X. 
Literature cited . 
. 264 
V. 
The Periderm . 
• 259 
Explanation of Plate . 
. 264 
VI. 
The Leaf Bases 
• 259 
I. Introduction. 
T HE specimen of a Medullosean stem which forms the subject of the 
present investigation was obtained from the colliery at Shore 
Littleborough in Lancashire, and is of Lower Coal Measure age, as are 
also the only other British species of the genus which have, up to the 
present, been described — namely M. anglica 1 and M. pusilla . 2 
The coal-ball in which the stem occurred was divided by Mr. J. Lomax 
into three parts : from the portion named 3 a series of six transverse 
sections was obtained (sections A-F 3 ), two slightly oblique transverse 
sections were available from part 2 (sections 8-9 4 ), and a transverse and 
two longitudinal sections were cut from piece 1 (sections 24 5 and 25-6 6 ). 
No information was obtained by which the distance between sections F and 
1 Scott, D. H. : On the Structure and Affinities of Fossil Plants from the Palaeozoic Rocks. 
III. On Medullosa anglica , a new Representative of the Cycadofilices. Phil. Trans. Roy. Soc., 
Ser. B, vol. cxci, 1899. (Referred to subsequently as M. anglica , loc. cit.) 
2 Scott, D. H. : Studies in Fossil Botany. Part II, 2nd ed., 1909; On Medullosa pusilla , 
Proc. Roy. Soc., Ser. B, vol. lxxxvii, pp. 221-8. 
3 University College, London, Collection, Catalogue nos. O. 800, O. 80/. 
4 „ „ „ „ „ nos. O. 80^, O. Soh. 
5 ,, ,, ,, ,, ,, no. O. 80 k. 
6 „ ,, „ ,, „ nos. O. 81 a, O. 81 b. 
[Annals of Botany, Vol. XXVIII. No. CX. April, 1914.] 
