Fagaceae , and its Bearing on the Affinities of the Group . 515 
axial prolongation, thus forming the greater part of the central column 
of the ovary, including the outer layers in which the ovular bundles run. 
The fruit of C. chrysophylla , like that of some of the Oaks, takes 
two years to develop. At the time of pollination, about the middle of 
July, ovules are present, but the archesporial cells are hardly distinguishable 
from the rest of the nucellar tissue, and the integuments are only beginning 
to appear. Little development takes place during the winter months, and 
fertilization does not occur in England till the latter half of the month 
of May. 
Cupule. 
There can be little doubt that in Castanopsis chrysophylla each valve 
of the cupule represents, not a bracteole covered with emergences, as 
— it 
b 
Fig. 3. Commencement of cupule valve, showing its growing-point (g.p.) and its position 
corresponding with that of the secondary flower of the dichasium. o.w., ovary wall of primary flower ; 
b. y bract of primary flower, x 45. 
Eichler suggests in his account of the cupule of other members of the 
Fagaceae, but one of the secondary axes of the dichasium, beset with 
