54 2 Worsdell . — Morphology of the ‘ Corona ’ of Narcissus. 
anther lobes having the colour and consistence of a corona, there were, 
in addition, two upwardly directed, broad, petaloid lobes (Fig. 2), so that in 
this case the corona was divided into two parts, exactly corresponding to 
the two parts into which the metamorphosed petal was divided, viz. upper 
anther lobes and lateral petaloid portions. In a third petal the basal 
anther lobes were partly polliniferous and partly corona-like. 
With these clear transitional structures before me, there can no longer 
be any doubt in my own mind as to the true morphological interpretation 
of the corona. Moreover, this could never have been arrived at without the 
aid of these teratological data. 
Fig. 1. Narcissus Pseudo-nar- 
cissus var. Staminoid petal show- 
ing ‘ corona * as a downward pro- 
longation of anther lobes. Letter- 
ing as in Fig. 2. 
Fig. 2. Ditto ; the extreme 
petaloid portion of ‘ corona ’ 
is upwardly directed, p. = 
petal ; an. — anther ; con. = 
connective : co. — corona. 
(Slightly schematized.) 
Fig. 3. Narcissus Pseudo- 
narcissusv&x. Perianth seg- 
ment ( p .) with itstwo-lobed 
corona segment (co.), show- 
ing transition to anther struc- 
ture. (Semi-diagrammatic.) 
But the point of chief interest has yet to be mentioned. In the flowers 
from whose structure Celakovsky deduced his remarkable conclusions, it was 
the petaloid stamens which exhibited the transitional conditions of the corona, 
and from which he further deduced the staminal origin of the perianth. In 
my own flowers it was the staminoid petals which exhibited the interesting 
transformations in the corona, which latter is thus seen to be changing 
back again to its original, primitive condition, viz. of basal anther lobes. 
Celakovsky’s view of corona-morphology becomes thus doubly supported, 
viz. by the occurrence of the same transformations taking place in both 
directions: progressively and retrogressively ! 
There is one further point of importance to be noted. In the change 
which is under way, whereby the three petals become partially transformed 
into stamens, the entire corona of the flower appears to become involved , the 
sepals being left without any trace of this structure ; Irom which it might 
be reasonably deduced that in Narcissus the normal corona is an outgrowth 
of the corolla only, and that the sepals, although equally derived from 
