Worsdell. ■ — The Morphology of the ‘Corona' of Narcissus. 543 
stamens possessing versatile anthers, no longer have, if they ever did have, 
any structure corresponding to the basal lobes of the anther. The petals of 
some genera of Amaryllidaceae, although derived by transformation of 
versatile anthers, are also devoid of any trace of a ligular outgrowth. On 
the other hand, Celakovsky, from a study of abnormal flowers of N. Tazetta , 
concluded that the calyx possessed a corona as well as the corolla, but that 
in some of the double flowers in which the stamens produced corona 
structures, that of the calyx was either greatly reduced or entirely suppressed. 
Hence it would be best, on the whole, to regard the calyx corona in the 
flowers I have been investigating as having also become suppressed as a 
result of the changes undergone by the corolla. 
Since writing the above note another abnormal flower, of the same 
species, has come into my hands, which shows almost equally important 
features. The corona has undergone complete dialysis, and each segment 
of the perianth bears a hilobed corona segment ; this character in itself 
represents, in my opinion, one step in the transition to the two basal lobes 
of the versatile antler (see above). But in the case of one of the six 
perianth lobes, the two lobes of its ventral corona segment each possessed 
an infolded flap on its mner margin , the two flaps being united below (i. e. 
towards the base of the corona segment) to form a short enation from the 
upper surface of the corona (Fig. 3). This structure, in my opinion, 
undoubtedly represents a still further step in the direction of transformation 
of the individualized corona segment into the basal lobes of the versatile 
anther, of which the perianth segment (/.) represents the apical portion. 
The following conclusions, therefore, are reached as a result of this 
investigation : 
1. Both sepals and petals, in Amaryllidaceae, have been derived by 
transformation of stamens in an originally achlamydeous flower. 
2. The corona, in this order, has been derived by petaloid transfor- 
mation of the upturned basal lobes of the versatile anthers, and subsequent 
fusion thereof to form a continuous rim. 
Kew, 
March 30, 1914. 
