652 
Campbell. — The Structure and A ffinities of 
whether it came from Sarawak or from Dutch Borneo. The plant was 
somewhat smaller than the specimens of M. Alidae from Bau, but it was 
assumed to be the same species. A further examination, however, has 
shown a number of differences, especially in the sporangium, and it should 
probably be considered specifically distinct. The plant at Buitenzorg is 
the type of Raciborski’s Angiopteris Smithii } but there is no question that 
it should be transferred to the genus Macroglossum , and would therefore 
become Macroglossum Smithii. 
The full-grown plant is very striking in appearance. From the short 
caudex arise the closely set, simply pinnate, massive leaves, which attain 
a length of nearly four metres. They stand almost vertical, and the 
appearance of the plant is very different from that of any species of 
Angiopteris > whose bipinnate leaves are much less closely set, and usually 
spread out widely from the caudex. The leaves are arranged spirally 
about the stem, which is apparently always radial in structure. The leaf- 
base is much enlarged and the stipules very conspicuous. The latter are 
united in front by a broad commissure (PI. XLVIII, Fig. c). The leaves 
are simply pinnate, the pinnae, which are closely set, being sometimes 
more than 50 cm. in length. They are very thick, with quite smooth 
margins in the adult plant. 
In general habit the plant more nearly resembles a large Danaea than 
it does Angiopteris. It also is superficially in habit like some of the large 
species of Zamia. This was remarked when the plant in the Buitenzorg 
gardens was compared with a Zamia growing not far away from it. 
Plate XLVIII, Fig. B shows the caudex of a small individual with 
the leaf bases and the thick, much-branched roots. 
The plants grew on a wooded bank above a small stream, and by 
examining the earth below the larger plants a number of prothalli were 
secured. As is usual with the Marattiaceae, the prothalli were most abundant 
where the earth had fallen away leaving freshly exposed surfaces. They 
are easily distinguished from the prothalli of ordinary Ferns by the more 
massive structure and the emergence of the primary leaf upon the upper 
surface of the gametophyte. Gametophytes of different ages were 
secured, some of unusually large size ; and a number of young sporophytes 
of different ages were also collected. 
The Gametophyte. 
As in all the Marattiales, the gametophyte of Macroglossum is a dark- 
green thallus with a very thick midrib which projects strongly on the 
lower surface (PL XLVI, Fig. 4). The wings of the thallus, also, are com- 
paratively thick and never composed of more than a single layer of cells, as 
1 Raciborski, M. : Uber einige tmbekannte Fame des Malayischen Archipels. Bull. Acad. d. 
Sciences, Cracovie, 1902. 
