Macrogtossu m Alidae, Copeland. 663 
leaf-traces. Several mucilage ducts (ml) were present in the neighbourhood 
of the vascular bundles, but none were seen in the cortical region. 
The Roots. 
The roots (Text-fig. 6) differ from the other Marattiaceae that have 
been examined in the complete ab- 
sence of mucilage ducts. There are 
numerous large tannin sacs, some of 
which approach the size of mucilage 
ducts, but the latter seem to be quite 
lacking. In the outer part of the 
cortex a few scattered sclerenchyma 
cells were seen, but there is no distinct 
ring of sclerenchyma such as occurs 
in the roots of Danaea. The stele 
of the root is much like that of other 
Marattiaceae, but the xylems are less 
distinct than is usually the case, and 
extend almost to the centre of the 
cylinder. The specimen figured had 
eleven xylems, but there is doubtless 
some variation in this respect. 
The Sporangia. 
The sorus in Macroglossum is very much longer than in any species of 
A ngiopteris, and much more nearly resembles that of A rchangiopteris. 
It may reach a length of 8 mm., and the number of sporangia in M. Alidae 
about 70. This is something more than half the maximum number in 
A rchangiopteris} The appearance of the sorus is very different from that 
of Angiopteris, as the individual sporangia are less evident, and they are 
very closely crowded, so that the elongated sorus, as we have seen, closely 
resembles a true synangium like that of Danaea. In reality, however, the 
sporangia are quite separate, as in Angiopteris. The resemblance of the 
sorus to the synangium of Danaea is increased by the development of an 
elevated ridge between the sori, which are thus partially sunk below the 
surface of the leaf. The sorus is surrounded by a conspicuous fringe of 
indusial hairs, which arise from the base of the sporangia and reach to the 
top. These indusial hairs, which branch repeatedly, are much more evident 
than in any species of Angiopteris , and are very similar to those found in 
A rchangiopteris. 
The sporangia (Text-fig. 7) also are more like those of A rchangiopteris 
than they are like those of Angiopteris. Owing to their position they are 
1 Christ, H., and Giesenhagen, K. : Pteridographische Notizen. Flora, lxxxvi, 1899. 
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Text-fig. 6. Section of root of an 
older sporophyte. x 16. 
