Macroglossu m Alidae , Copeland. 667 
growing in the Botanical Gardens at Buitenzorg, Java, but of unknown 
origin. 
2. The full-grown plant approaches in size the larger species of 
Angiopteris , the leaves may reach a length of 4 metres. The plant differs 
much in appearance from Angiopteris , the leaves being more numerous and 
upright, and simply pinnate. The very much elongated and partially 
immersed sori are also strikingly different from those of Angiopteris. 
3. The gametophyte is much like that of Angiopteris. It may reach 
a length of 3 centimetres. Branching is not uncommon, either by ad- 
ventitious buds, or occasionally by a dichotomy of the growing point. 
4. The antheridia occur both on the dorsal and ventral surfaces. They 
are of the usual type, but the number of sperm-cells is probably greater 
than in any other Marattiaceae. The spermatozoids are relatively small 
and less elongated than those of Angiopteris . 
5. The archegonia are much like those of Angiopteris. 
6. The embryo develops a conspicuous suspensor and thus resembles 
the embryo of Danaea. 
7. The later development of the embryo seems to correspond to that 
of the other Marattiaceae. The apical cell of the stem is not always clearly 
demonstrable. 
8. As in the other Marattiaceae, no true cauline stele is developed in 
the young sporophyte, and the vascular system of the axis is composed 
entirely of leaf-traces. 
9. The structure of the axis in the young sporophyte is to a certain 
extent intermediate between that of Danaea and Angiopteris. 
10. The cotyledon is much like that of Angiopteris ; the first leaves, 
about ten, remain simple. Stipules first appear in the third leaf. The 
first leaf-trace shows a collateral structure. 
11. The primary root is usually diarch. A single root is formed for 
each of the early leaves. The primary root has a single initial cell. 
12. The leaf of the adult sporophyte differs from that of Angiopteris 
in its simply pinnate form, as well as in the anatomy of the leaflets. The 
latter show a conspicuous mucilage duct below the sorus, and a better 
developed palisade tissue. 
13. The sporangia of Macroglossum , especially M. Alidae , are smaller 
than those of Angiopteris , and very much more numerous. The sorus is 
much more compact, and partially sunk in a trough formed by elevated 
ridges between the sori, while in Angiopteris no such ridges are present. 
The annulus is very slightly developed in Macroglossum , and the indusial 
hairs very much more conspicuous than in Angiopteris. 
14. The roots of the adult plant have no mucilage ducts, thus differing 
from the other Marattiaceae that have been described. 
15. Macroglossum should undoubtedly be placed in the Angiopteridiae, 
