The Anatomy of some Xerophilous Species of 
Cheilanthes and Pellaea. 
■ BY 
A. S. MARSH, B.A. 
With eleven Figures in the Text. 
Material. 
HE species examined were all collected by Mr. A. G. Tatisley either 
X in America or in Algeria, and to him I am indebted for my material, 
and for his very great assistance throughout the investigation. 
The plants were Cheilanthes per sica, gathered in 1910 at Djebel 
Murdjadjo, Oran ; Cheilanthes Fendleri y from Santa Catalina, California ; 
Ch. lanuginosa , from Manitou, California ; Ch. gracillima, from Crater Lake, 
Oregon ; and Pellaea andromedae folia , from Santa Catalina, California. The 
four last-mentioned species were all obtained in the Western United States 
of America in the summer of 1913. 
For purposes of comparison, Pellaea f ale ata, a New Zealand mesophytic 
Fern, was also investigated to some extent. 
External Morphology and Xerophilous Adaptations 
in the Leaves. 
The fronds of all four species show well-marked xerophilous character- 
istics, which have been concisely expressed as follows in the * Synopsis 
Filicum ’ : 
‘ Cheilanthes lanuginosa , Nuttall. Frond bipin natifid, rachis tomentose, 
texture subcoriaceous ; upper surface grass green, slightly tomentose ; 
lower surface densely matted with a coat of pale brown woolly tomentum ; 
the margin of the segment much incurved. 
‘ Cheilanthes per sica, Mett. ( — Ch. Szovitzii , Fisch. & Meyer). Frond 
bipinnatifid, rachis scaly and tomentose, texture subcoriaceous ; upper 
surface green, slightly tomentose ; lower surface densely matted with 
a coat of pale brown woolly hairs ; the margin of the segment much 
incurved. 
[Annals of Botany, Vol. XXVIII. No. CXII. October 1914.] 
