7° 2 
Thomas . — Seedling Anatomy of 
Anemoneae. 
Anemone coronaria . This species has a marked cotyledonary tube in 
which two small collateral strands, corresponding to the midribs of the 
respective cotyledons, face each other across the central cavity. Imme- 
diately below the cotyledonary node the strands close into the centre of 
the axis through a sharp bend, so that the tissues are almost longitudinal in 
the transverse section, and this may be sometimes more marked on one 
side than on the other. As the strands enter the axis, the readjustments 
take place which produce a diarch root very near the top of the hypocotyl. 
These readjustments are obscured by the fact that, to a large extent, they 
take place in the bend, but in the section shown in PL L, Fig. 3, the bifurca- 
tion of the phloem can be seen very plainly in the strand to the south. 
The aggregation of the phloem in two alternating patches is the root 
character which appears highest in the hypocotyl, while the absence of pith 
and formation of a definite xylem plate with exarch protoxylem groups 
are characters only to be found at a lower level. Although each cotyle- 
donary bundle consists of only three or four xylem elements, one or two 
of these are usually secondary in formation, judging from the radial 
arrangement found. 
Anemone ftdgens forms a cotyledonary tube of considerable length, 
which in the upper part becomes a solid cylindrical structure through the 
interlocking of cells of the cavity (Text-fig. 1). The cotyledon strands 
pass more steeply into the hypocotyl, so that the transition can be followed 
with greater ease than in A. coronaria. 
Anemone rivularis is very similar to A. ftdgens. 
Anemone sylvestris . In this a very short cotyledonary tube or sheath 
is found, which in some individuals is asymmetrical through union of the 
cotyledons at a slightly higher level on the south side away from the 
first leaf, and through displacement of the cotyledonary strands to this 
side. 
The transition features are similar to those of A. coronaria . 
Anemone mrginiana. In this species the cotyledons fuse quite at the 
base as i n A. sylvestris (Text-fig. 2). In this region the cotyledon strands 
are very slightly double groups. The features of transition are like those 
of the above-described species. 
Anemone pidsatilla . There is a very slight cotyledonary tube at the 
base, in which the cotyledon bundles show a double structure. 
In one individual the most remarkable asymmetry was found, resulting 
in the formation of a diarch root, one of the poles of which is continuous 
with the vascular system of the first plumular leaf, while the other is 
continuous with the two asymmetrically placed cotyledonary strands. 
Anemone vitifolia also possesses a short cotyledonary tube, which is 
