xo6 Campbell. — The Embryo-Sac of Peperomia. 
was not the same as the one cultivated under the same name 
at Kew. 
The behaviour of the other nuclei, at the apex of the sac, 
is not always the same. In some cases observed, where there 
were two of these, they were arranged with reference to the 
egg-cell in such a manner as to suggest the typical angio- 
spermous egg-apparatus. Johnson 1 states that he always 
found a single conspicuous synergid, which after fertilization 
developed a cell-wall and persisted until the embryo was 
nearly full-grown. The writer found, in a good many cases, 
preparations which showed an appearance not unlike that de- 
scribed, but it was impossible to see in what essential respect 
the so-called synergid was different from the other nuclei 
which do not form part of the group already referred to. 
These also develop a membrane about them, and are often 
quite as conspicuous as the alleged synergid (Fig. 6, B, x) 
which we are inclined to think is nothing more than one of 
these nuclei with its accompanying cytoplasm, which simply 
is close to the egg. As there may be two, or even three of 
these cells in the upper part of the embryo-sac, and they 
take no part in the conduction of the generative nucleus 
to the egg, it is very questionable whether they can be 
properly spoken of as synergids. 
At the time the pollen-tube reaches the embryo-sac, the 
egg-cell, which may be slightly pushed to one side, appears 
as a flattened body, and close to it there are from one to 
three nuclei, with a more or less definite aggregation of 
cytoplasm about them, and perhaps to be considered as the 
morphological equivalents of the synergidae of the typical 
embryo-sac. They take no part, however, in the process 
of fecundation. 
The group of (usually) eight nuclei already referred to 
(Figs. 2, 3), may at the time of fertilization occupy the base of 
the sac, or these nuclei may be aggregated at a point on one 
side, or even close to the egg-cell. These, as Johnson has 
1 loc. cit., p. 2. 
