246 Bower . — Imperfect Sporangia in certain 
the further development important differences from other 
European species arise, and the embryo in its later form 
resembles that of L. Phlegmaria. The foot remains small 
as a conical continuation of the large suspensor : there is no 
protocorm, nor are there any protophylls of the Phylloglosskm 
type: the first leaves are green, and resemble the foliage 
leaves of the mature plant of L. Selago : these are carried up 
to the surface of the soil, whatever the position of the pro- 
thallus — buried or at the surface — by a proportional inter- 
calary growth in the hypocotyl, which is directly continuous 
with the foot 1 2 : the first root is formed near its base. The first 
branching of the axis occurs in plants which have about six 
or seven developed leaves : ‘ after the second branching these 
seedlings may produce spores, while in other native species 
only quite old plants proceed to spore-development V These 
characters, together with those of the sporophyte, are recog- 
nized by Bruchmann as showing that Z« Selago is not near to 
any of the species native in Germany 3 . 
Perhaps the most important of these facts is the early 
appearance of the sporangia on the seedling of Z. Selago. 
It shows, in the only species of the Selago group yet examined 
embryologically, that early spore-production goes along with 
the undifferentiated condition of the whole plant. The stele 
also of the mature plant is of simpler structure, and probably 
more primitive type than those of other species whose 
embryology and anatomy are both known. These characters 
together indicate the primitive nature of the group, and 
suggest the further question, is the embryology also of 
Z. Selago to be regarded as a primitive type, compared with 
that of other Lycopods, and if so what will the effect be on 
our views regarding Phylloglossum and the theory of the 
protocorm ? 
Before Treub’s paper in which this term was introduced, 
and the theory of the protocorm enunciated 4 , the ‘ embryonic 
1 Bruchmann, loc. cit., PI. VI, Figs. 1-26, and PI. VII, Figs. 41-43. 
2 lb., p. 100, and PI. VI, Figs. 26-27. 3 Loc. cit., p. 102. 
4 Buitenzorg, Annals, vol. viii, p. 1. 
