Boodle . — Anatomy of the Schizaeaceae. 371 
rhizome, it has a nearly longitudinal course until after passing 
through the phloem region of the rhizome. It then curves 
out more sharply. In Lygodium japonicum , at the junction 
of the xylem of the root with that of the stem, a few branched 
tracheides occur, but they differ from those at the node, in 
that their arms are scarcely divergent. This depends on 
the angle at which the root-stele separates. In all the 
species ‘ root-tracheides ’ are differentiated at the periphery 
of the stele a short distance before they ‘ pass out ’ to the 
root. 
Lygodium, seedling. 
The seedling 1 of Lygodium japonicum may now be de- 
scribed. The first leaf is simple and palmatipartite ; the 
second leaf has its lamina dichotomously divided into two 
palmatipartite segments. In the next few leaves the petiole 
forks into two at the top with one pinnately lobed pinna on 
each fork. The later leaves have the growth of the rachis 
continued above the two pinnae as a circinate tendril-like 
projection or a bud. The appearance of the rhizome in 
oldish seedlings suggests that, after the first dichotomy of 
the rhizome, one branch does not develop far, and that growth 
is continued by forking of the other branch. The roots show 
distichous arrangement. 
The structure of the seedling was examined by means of 
microtome-series. Several series were made, and showed 
differences in detail, but agreement in the main characters. 
The primary root is diarch. In the region of the foot the 
root-stele curves sharply (nearly horizontally) inwards to- 
wards the prothallus. Fig. 8 shows the arrangement of the 
foot (/.) and prothallus (p.) at the point where the root-stele, 
whose diarch xylem is shown diagrammatically at x., is just 
beginning to curve inwards. After curving in, it turns 
upwards so as to become vertical again, and generally by 
that time has lost the distinctive characters of root structure, 
1 ‘ Sporeling * is of course the more reasonable term, but ‘ seedling ’ appears 
customary. 
Bba 
