Boodle,-— Anatomy of the Schizaeaceae. 413 
be pointed out as rather characteristic features in the Schizae- 
aceae. 
Four types of stele are represented among the Schizaea- 
ceae: — the solid stele in Lygodium , the medullated stele in 
Schizaea , solenostely in some species of Anemia , and dialystely 
in other species of Anemia and in Mohria . With regard to 
Schizaea , as the transitional region of seedlings has not 
yet been specially examined, it cannot be said whether any 
different type of structure occurs at an early stage in the 
ontogeny, as seen by Jeffrey in some Angiosperms. De Bary 
(’ 77 , p. 344) agrees with Russow’s view that the central tissue 
within the xylem of Schizaea is not to be regarded as a pith, 
but as parenchyma belonging to the stele. But as pith is not 
a sharply characterized term there is no harm in applying 
it to this tissue, especially if one inclines to the view that 
phylogenetically pith is stelar tissue. A structure is found 
in Anemia coriacea, which though resembling solenostely may 
differ in the behaviour of the endodermis at the node. 
The petiole-bundle is concentric in Lygodium , collateral in 
Schizaea , and roughly speaking bicollateral in Anemia and 
Mohria. 
The structure of Anemia and Mohria must be compared, 
as Christ (’ 97 , p. 352) regards the external characters of the 
latter as so anomalous for the Order. Mohria resembles 
Anemia Phyllitidis completely in the type of its stelar 
system, and agrees roughly with other species of Anemia , 
as to the dimensions of the parts. The petiolar bundle in 
Mohria is of the type found in Anemia. There are no fibres 
in the bundle of Mohria , but that is also true for certain 
species of Anemia, e. g. A. Dregeana. Mohria possesses 
glandular hairs exactly similar to those of Anemia , but differs 
in possessing paleae. An approach to palea-formation occurs 
however in Lygodium , therefore this character is not of great 
importance. Again, the sclerotic group accompanying the 
petiole-bundle of Mohria is suggested by the thick-walled 
elements which occur in a corresponding position with regard 
to the bundle at the base of the petiole of Anemia Phyllitidis. 
