Lew ton- Brain.- — Cor dy ceps ophioglossoides (Ehrhi). 527 
* by Reess, I thought it advisable to give a short account of the 
mycorhiza-structure, so far as could be made out from the 
material at my disposal. 
Reess describes the mycorhiza as being ectotrophic and 
says, at all events for the young stages, that the Fungus 
extends into the root only a distance equal to the thickness 
of the fungus-sheath outside. He describes these internal 
hyphae as being intercellular, and as giving rise to peculiar 
mulberry-like ingrowths into the cell-cavity of the cortical 
cells. He is in doubt as to whether these structures are 
haustoria, or merely hypertrophies of the cell-wall caused 
by the action of the intercellular hyphae ; he seems on the 
whole to favour the former view, at any rate in some cases, 
while in others the growths appear to show no lumen and give 
the reactions of ordinary parenchyma-membranes, while their 
connexion with the intercellular hyphae is doubtful. These 
structures are described as being conspicuous and of frequent 
occurrence. 
My observations show that Elaphomyces , as a mycorhiza 
Fungus, is rather variable in its structure. 
The structure most frequently found shows in the young 
stage the ordinary fungus-sheath on the outside of the root, 
and this is continuous with an intercellular network (Fig. 25) 
between the cells of the root-cortex. The network in surface 
view has the appearance of a reticulate thickening of the cell- 
walls. This structure is that described by Frank 1 , for the 
mycorhiza on the roots of Cupuliferae, but in his case only 
the outermost layer of cells of the root was concerned. In 
my sections the whole of the root-cortex was seen to be 
penetrated in this way by the Fungus (Fig. 24). 
One very characteristic feature of this mycorhiza is the 
action of the Fungus on the cortical cell-contents. These are 
transformed into (apparently) slimy, deeply staining masses, 
which contain fat (Figs. 25, 27, 28, 29). This change takes 
place right through the cortex of the root. Sometimes a 
large, more or less disorganized nucleus is present in the mass. 
I Bot. Zeit., 18S5. 
