Comparative Anatomy of the Hymeno- 
phyllaceae, Schizaeaceae and Gleicheniaceae. 
Ill, On the Anatomy of the Gleicheniaceae 1 . 
BY 
L. A. BOODLE, F.L.S. 
With Plates XXXVIII and XXXIX. 
T HE fullest account of the anatomy of the Gleicheniaceae 
previously published is that given by Poirault (’98), but 
although he enters into a careful description of the petiole 
and node in different species of Gleichenia , the structural 
characters of the Order have in some respects been less 
completely dealt with than was the case with the Hymeno- 
phyllaceae and Schizaeaceae, which formed the subject of 
two monographs by Prantl (75 and ’81). 
The range of diversity in the structure of the rhizome is 
less in the Gleicheniaceae than in the Schizaeaceae. In the 
latter Order four main types of stelar structure were recog- 
nized, while in the Gleicheniaceae only three are found, as will 
be described below. No case of dialystely has so far been 
discovered. The homologies of the three types of structure 
which occur in the Gleicheniaceae is an interesting question, 
but, although it is as well to discuss it, there does not 
1 From the Jodrell Laboratory, Royal Botanic Gardens, Kew. 
[Annals of Botany, Vol. XV. No. LX, December, 1901. J 
