On the Structure and Affinities of Sutcliffia, 
in the Light of a Newly Discovered Specimen. 
BY 
E. DE FRAINE, D.Sc., F.L.S. 
University of London ; University College. 
With Plates XCI and XCII and nineteen Figures in the Text. 
Contents. 
I. 
Introduction . 
PAGE 
1031 
V. 
The Ground Tissue and the 
PAGE 
IT. 
General Structure 
1033 
Secondary Cortex 
1051 
III. 
Course of the ‘Meristeles’ 
1037 
VI. 
T he Extrafascicular Strands 
1053 
IV. 
Histology : — 
VII. 
Attribution of the Specimen 
1056 
1. Structure of the Wood 
io 43 
VIII. 
Affinities . 
1058 
2. Structure of the Phloem 
1046 
IX. 
Summary 
1063 
3. Structure of the Leaf-trace 
Bundles . 
io 47 
Literature cited . 
Explanation of Plates . 
1064 
1066 
I. Introduction. 
T HE fossil plant described in the present paper was obtained from the 
colliery at Dearnley near Littleborough in Lancashire in 1910, and is 
of Lower Coal Measure age. It occurred in a nodule from the roof of the 
workings, as is shown by the presence of Goniatite shells scattered through 
the sections, and probably was obtained from the same seam as the type 
specimen, Sutcliffia insignis , which came from a roof-nodule from the 
adjacent mine at Shore Littleborough. 1 The specimen, which was about 
10 inches long, was cut by Mr. James Lomax, and thanks are due to the 
‘ Committee of British Fossil Plants ’ of the British Association for the 
Advancement of Science for a grant which partially defrayed the cost 
entailed in cutting the specimen. In September, I9ii,the series of fifty- 
eight transverse and nineteen longitudinal sections, labelled c new Medullosa, 
probably Sutcliffian type \ was handed over to me by Professor F. W. 
Oliver, F.R.S., for investigation. I cannot allow this opportunity to pass 
without gratefully acknowledging the never-failing help and advice which 
1 Scott, D. H. : IV. On Sutcliffia insignis , a New Type of Medulloseae from the Lower Coal 
Measures. Trans. Linn. Soc. London, vol. vii, Pt. 4, ser. 2. Sept. 1906. 
Annals of Botany, Vol. XXVI. No. CIV. October, 1912.] 
V 
