356 
Essed . — The Panama Disease. II. 
which one or more plasmic globules, described before as spore-initials, are 
seen to form. In maturing they assume a dark violet colour and are liberated 
as large chlamydospores, or rather chlamydosporangia, measuring 80-120 
feet (see PI. XXIX, Fig. 3). They do not differ at all from the bodies which 
were mentioned in my first paper under the name of giant chlamydospores. 
In germinating the spore coat was seen to burst, allowing the germ tube to 
emerge. This germ tube consisted of one or more club-shaped cells ; from 
the peculiar way in which they adhere to each other I infer that each cell 
is a separate germ tube derived from a special spore-initial, and that the 
adherence is only due to compression caused by growth in a limited space 
(see Fig. 4). The colour of the germ tube is dark, producing a mycelium 
which at first is also dark, but gradually becomes hyaline (see under Pure 
cultures ’ in first paper). Besides the mode of oidium-formation mentioned in 
my first paper, I found some hyphae of the mycelium treated of above forming 
chains of club-shaped bodies (i. e. oidia). Some of these oidia passed into 
chlamydospores. Sickle-shaped conidia on parts of the same mycelium 
also underwent the same transformation (see Fig. 3). The chlamydospore- 
fructification is undoubtedly the most prominent feature of the reproductive 
habit of this fungus. 
Haastoria. They arise as little knob-like excrescences, lateral or 
terminal, with opaque contents protected by a thin membrane. They 
grow out into flat saucer-like structures, or assume a funnel or a spoon 
shape, or become polypoid or tassel-like (see Fig. 5) ; they appear to 
assimilate food with the aid of their secretion, the turbidity of the proto- 
plasm and the gummy degeneration of the walls of the cells, in which they 
arise, giving support to this assumption. The hyphae from which they 
arise become irregularly distended and turn in the end to pegmatia. The 
haustoria themselves may be wholly or partially transformed into myco- 
poroma. This takes place at the close of the parasitic mode of life of the 
fungus. 
Pycnidia. Under this name were mentioned spore-masses found in 
the disintegrated tissues of the decaying rhizome and leaves. In fact they 
are better termed pseudo-pycnidia, since they arise in pre-existing cavities 
due to rupturing in consequence of contraction in the putrifying tissues. 
The shape is spheroidal or irregular. From all directions fertile hyphae 
enter the cavities and mainly crowd together in the lining cells. From this 
crowding together a felted mass ensues, out of which arise the mycoporo- 
matic lining of the cavities and the huge number of chlamydospores filling 
them. Sometimes multicellular conidia are found among the chlamydo- 
spores ; they are probably abstricted from hyphae, traversing the cavities 
before the chlamydospore-formation began (see Fig. 6). Considering the 
fact that some of these pseudo-pycnidia were mainly filled with conidia, and 
some conidia were found in a state of transformation into chlamydospores, 
